Alice R. Millikin,^1,*,** Drew R. Davis,^2,3 Donald J. Brown,^1,4,*** Sarah K. Woodley,⁵ Stephanie Coster,⁶ Amy Welsh,¹ Jacob L. Kerby,⁷ James T. Anderson<sup>8

1</sup>School of Natural Resources, West Virginia University, 1145 Evansdale Drive, Morgantown, West Virginia 26506, USA
²School of Earth, Environmental, and Marine Sciences, The University of Texas Rio Grande Valley, 1 W University Boulevard, Brownsville, Texas 78520, USA
³Biodiversity Collections, Department of Integrative Biology, The University of Texas at Austin, 10100 Burnet Drive, PRC 176-R4000, Austin, Texas 78758, USA
⁴USDA Forest Service, Northern Research Station, PO Box 404, Parsons, West Virginia 26287, USA
⁵Department of Biological Sciences, Duquesne University, 600 Forbes Avenue, Pittsburgh, Pennsylvania 15282, USA
⁶Department of Biology, Randolph-Macon College, 114 College Avenue, Ashland, Virginia 23005, USA
⁷Department of Biology, University of South Dakota, 414 E Clark Street, Vermillion, South Dakota 57069, USA
⁸James C. Kennedy Waterfowl and Wetlands Conservation Center, Belle W. Baruch Institute of Coastal Ecology and Forest Science, Clemson University, PO Box 596, Georgetown, South Carolina 29442, USA

^*Current address: Natural Resources Conservation Service, 2322B Goddard Parkway Salisbury, Maryland 21801, USA
^**Corresponding author (email: alicemillikin@gmail.com)
^***Current address: USDA Forest Service, Pacific Northwest Research Station, 42218 NE Yale Bridge Road, Amboy, Washington 98601, USA