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Additional Information

Competing Interests

There are no competing interests

Author Contributions

Heather Paxton conceived and designed the experiments, performed the experiments, analyzed the data, contributed reagents/materials/analysis tools, wrote the paper, prepared figures and/or tables, reviewed drafts of the paper.

Peter G Tickle conceived and designed the experiments, performed the experiments, analyzed the data, contributed reagents/materials/analysis tools, prepared figures and/or tables, reviewed drafts of the paper.

Jeffery W Rankin conceived and designed the experiments, performed the experiments, reviewed drafts of the paper.

Jonathan R Codd conceived and designed the experiments, reviewed drafts of the paper.

John R Hutchinson conceived and designed the experiments, reviewed drafts of the paper.

Animal Ethics

The following information was supplied relating to ethical approvals (i.e., approving body and any reference numbers):

The Royal Veterinary College ethics committee

Grant Disclosures

The following grant information was disclosed by the authors:

BBSRC BB/I0211161/1

Funding

This work was supported by a BBSRC grant (BB/I021116/1) to JRH and JRC. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript.


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