CC BY
340E30PM REVIEW ARTICLES
LIST OF SEVERAL FAMILIES OF DIPTERA FROM THE VOLGA UPLAND AND THE OKA-DON LOWLAND
A.B. Ruchin1*, N.E. Vikhrev2, D.I. Gavryushin2, M.N. Esin1
1 Joint Directorate of the Mordovia State Nature Reserve and Smolny National Park, Russia 2Zoological Museum of Moscow University, Russia e-mail: l*ruchin.alexander@gmail.com, 2nikita6510@ya.ru
Received: 01.12.2022. Revised: 15.01.2023. Accepted: 01.02.2023.
Diptera is one of the most species-diverse insect orders. The paper presents data on the biodiversity of Diptera from 9 families (Tipulidae, Limoniidae, Pediciidae, Mycetophilidae, Keroplatidae, Ptychopteridae, Muscidae, Fanniidae, Sciomyzidae) inhabiting 14 regions within the Volga upland and Oka-Don lowland. The most studied fauna is in the Republic of Mordovia (399 species). The fauna of these families is least known from the Republic of Tatarstan, Lipetsk Region, Nizhny Novgorod Region, and Vladimir Region (9, 19, 20, and 21 species, respectively). A total of 545 species are known from 9 families of Diptera in the large macroregion. The largest number of species includes the families Muscidae (190) and Limoniidae (102). The biodiversity is least known in Ptychopteridae (4 species), Pediciidae (6 species) and Keroplatidae (10 species). Key words: Tipulidae, Limoniidae, Pediciidae, Mycetophilidae, Keroplatidae, Ptychopteridae, Muscidae, Fanniidae, Sciomyzidae, European Russia
https://dx.doi.org/10.24412/cl-31646-2686-7117-2023-32-5-67
Introduction
Diptera is one of the most species-diverse orders among the holometabolous insects. According to Pape et al. (2011), it has approximately more than 160 000 described species. However, new species are constantly being described from different parts of the world (Nedeljkovic et al., 2015; Maslova et al., 2020; Grichanov, 2020, 2021; Burington, 2022; Mondal et al., 2022; Moravvej et al., 2022; Yang et al., 2022; and many others). Diptera is traditionally divided into two suborders: Nematocera and Brachycera (Pape et al., 2011), whose degree of knowledge is very uneven in different parts of the world. For example, there are up-to-date checklists of individual Diptera families from different countries (Jezek et al., 2018; Korkmaz et al., 2021; El-Hawagry et al., 2022; Kettani et al., 2022; Mukherjee et al., 2022).
There is a large variety of environments where Diptera larvae and imago occur. For example, aquatic Diptera are found in many shallow streams and puddles, in turbulent waterfalls, in tree hollows, in glacial waters and hot springs. They have been found at altitudes up to 5600 m in the Himalayas and at depths of more than 1000 m in Lake Baikal. They are also the only free-living insects that have colonised aquatic environments on absolutely all continents (Adler & Courtney, 2019). Many
Diptera species are important bioindicators of ecosystem state (Maritano & Sommaggio 2020; Youbi et al., 2020; Lychkovskaya et al., 2022). A large number of species and groups are involved in pollination, herbivory, predation, decomposition and other ecosystem processes at different stages of the life cycle. Imagoes feed on pollen and nectar of Apiaceae, Asteraceae, Ranunculaceae and Rosaceae and others (Rotheray & Gilbert, 2011; Barkalov & Khruleva, 2021; Villa et al., 2021). Phytophage larvae feed on underground and above-ground parts of living plants (Ricarte et al., 2017; Souba-Dols et al., 2020). Phytophagous larvae feed on a wide range of insects, both in the understorey and above-ground parts of plants (Rojo et al., 2003; Chursina & Ruchin, 2018; Astakhov et al., 2019). Saprophagous larvae are filter feeders of bacteria and/or detritus found in decomposing materials of both animal and plant origin (Krivosheina & Krivosheina, 2019; Rotheray, 2019); within saprophages, those larvae that depend on dead wood or decomposing materials associated with tree parts are saproxial (Rotheray et al., 2001; Polevoi, 2021). This publication presents an overview of the diptera fauna of a significant macroregion in terms of area, occupying the Volga Upland and the Oka-Don Lowland.
Material and Methods
Study area
The Oka-Don Lowland lies in the European part of Russia from the Oka River (Meshchera Lowland) in the north to the Kalach Upland in the south. In the south, the plain becomes progressively narrower and it curves towards the mouths of the Khoper and Medveditsa Rivers (Volgograd Region). It is situated within the East European Plain, between the Central Russian and Volga Uplands. The relief is characterized by vast flat areas with an absolute height of 150-180 m, which alternate with wide valleys of large rivers, stretching almost meridionally (the Don, Voronezh, Bityug, Khoper, Medveditsa, Tsna, etc.). With few exceptions, only steep and high slopes of the major river valleys are intersected by gullies. Large rivers are cut into watersheds to a depth of 25-50 m. Interfluves and gully intervals near valleys and gullies are gently sloping, while the rest of the territory is a flat plain. Swampy limestone depressions are not uncommon in the northern parts of the plain. The climate is continental with moderately cold winters and warm summers, with a pronounced continental increase compared with the Srednerusskaya Upland, situated to the west. Summers are drier and hotter and winters are colder. The Ryazan Region, Penza Region, Volgograd Region, Voronezh Region, Tambov Region, Lipetsk Region, Vladimir Region and the Republic of Mordovia are located within this lowland area.
The Volga Upland lies on the right bank of the Volga River from Nizhny Novgorod in the north to Volgograd in the south. Heights of 150-180 m prevail in the northern part of the upland. To the south, they increase and reach 200 m (in the upper reaches of the Sura River and in the Zhiguli Mountains more than 300 m). In the latitude of the Zhiguli Mountains, the upland stretches over 500 km from east to west and in the south, near Volgograd, its width does not exceed 60 km. The Volga Upland stretches in a meridional direction from north to south for about 1000 km and therefore
falls into several climatic zones (Milkov, 1953). The upland is composed of relatively easily erodible rocks: sandstones, clays, marl, chalk and opoka. In the areas of the so-called mountains (Zhiguli, Khvalynsk etc.) harder and more ancient sediments -limestone and sandstone - are outcropped. The territory is heavily ploughed up. The climate is continental, often with warm and hot summers and cold winters. There are often strong winds. There is the following pattern: as one moves from northwest to southeast, the continentality of the climate increases, as eastern and southeastern influences increase and western and northwestern ones weaken. However, in some areas this pattern is disrupted by local conditions - upland topography and vegetation. The Nizhny Novgorod Region, Republic of Mordovia, Chuvash Republic, Republic of Tatarstan, Penza Region, Ulyanovsk Region, Samara Region, Saratov Region and Volgograd Region are located within this upland area (Fig.).
Making a list
The families in the list are listed in a systematic order. Names of genera and species are listed alphabetically. Our list consists of previously published data and newly reported records. We did not include in several clearly erroneous reports from previous publications. For newly reported records, the label data (locality, date, collector) is given. Geographical coordinates are given in the Decimal Degrees format.
The list of regions and their short names are indicated as follows:
Chuvashia - Chuvash Republic;
Lipetsk - Lipetsk region;
Mordovia - Republic of Mordovia;
Nizhny Novgorod - Nizhny Novgorod region;
Penza - Penza region;
Ryazan - Ryazan region;
Samara - Samara region;
Saratov - Saratov region;
Tambov - Tambov region;
Tatarstan - Republic of Tatarstan;
Ulyanovsk - Ulyanovsk region;
Vladimir - Vladimir region
Volgograd - Volgograd region;
Voronezh - Voronezh region;
The specimens examined in this study are deposited in the following institutions:
Mordovia State Nature Reserve, Republic of Mordovia, Pushta village (those from M. Esin, A. Ruchin, G. Semishin);
ZIN - Zoological Institute, St-Petersburg, Russia;
ZMUM - Zoological Museum of Moscow University, Russia (those collected by N. Vikhrev, K. Tomkovich and M. Yanbulat).
When using a very popular in Russia term "cordon" we mean a lodge on the territory of a nature reserve where rangers or other staff may live. Abbreviations: SNR - State Nature Reserve, vill. - village.
Results
List of species
SUBORDER NEMATOCERA TIPULIDAE
Ctenophora fastuosa Loew, 1871
Chuvashia (Paramonov, 2018a). Mordovia (Ruchin & Pilipenko, 2015). Saratov (Paramonov, 2019).
Ctenophora festiva Meigen, 1804
Saratov (Paramonov, 2015).
Ctenophora flaveolata (Fabricius, 1794)
Chuvashia (Paramonov, 2018a).
Ctenophora guttata Meigen, 1818
Samara (Rosenberg, 2007). Ulyanovsk (Paramonov, 2006).
Dictenidia bimaculata (Linnaeus, 1760)
Chuvashia (Volkova, 1934; Paramonov, 2016; Paramonov, 2017a). Mordovia (Ruchin & Pilipenko, 2015; Pilipenko et al., 2020). Samara (Rosenberg, 2007). Saratov (Paramonov, 2015). Ulyanovsk (Paramonov, 2006).
Nephrotoma aculeata (Loew, 1871)
Chuvashia (Paramonov, 2018a).
Mordovia (Ruchin & Pilipenko, 2015; Pilipenko et al., 2020).
Nephrotoma analis (Schummel, 1833)
Chuvashia (Paramonov, 2018a).
Mordovia (Ruchin & Pilipenko, 2015; Pilipenko et al., 2020).
Nephrotoma appendiculata (Pierre, 1919)
Chuvashia (Paramonov, 2016; Paramonov, 2017a). Mordovia (Ruchin & Pilipenko, 2015; Pilipenko et al., 2020). Ulyanovsk (Paramonov, 2006).
Nephrotoma cornicina (Linnaeus, 1758)
Chuvashia (Volkova, 1934; Paramonov, 2016; Paramonov, 2017a). Mordovia (Plavilshchikov, 1964; Ruchin & Pilipenko, 2015; Pilipenko et al.,
2020).
Ulyanovsk (Paramonov, 2006).
Nephrotoma crocata (Linnaeus, 1758)
Chuvashia (Volkova, 1934; Paramonov, 2016; Paramonov, 2017a).
Mordovia (Ruchin & Pilipenko, 2015).
Samara (Rosenberg, 2007).
Ulyanovsk (Paramonov, 2006).
Voronezh (Negrobov, 2005).
Nephrotoma croceiventris lindneri (Mannheims, 1951)
Ulyanovsk (Paramonov, 2006).
Nephrotoma dorsalis (Fabricius, 1781)
Chuvashia (Paramonov, 2017a). Mordovia (Pilipenko et al., 2020).
Nephrotoma lamellata (Riedel, 1910)
Chuvashia (Paramonov, 2017a). Mordovia (Pilipenko et al., 2020).
Nephrotoma lunulicornis (Schummel, 1833)
Ulyanovsk (Paramonov, 2006).
Nephrotoma pratensis (Linnaeus, 1758)
Mordovia (Ruchin & Pilipenko, 2015; Pilipenko et al., 2020). Ulyanovsk (Paramonov, 2006). Voronezh (Negrobov, 2005).
Nephrotoma quadrifaria (Meigen, 1804)
Chuvashia (Paramonov, 2018a).
Nephrotoma quadristriata (Schummel, 1833)
Mordovia (Pilipenko et al., 2020).
Nephrotoma rossica (Riedel, 1910)
Chuvashia (Paramonov, 2017a). Ulyanovsk (Paramonov, 2006).
Nephrotoma scalaris (Meigen, 1818)
Chuvashia (Egorov & Podshivalina, 2014; Paramonov, 2016; Paramonov, 2017a). Mordovia (Pilipenko et al., 2020). Ulyanovsk (Paramonov, 2006). Saratov (Paramonov, 2019).
Nephrotoma scurra (Meigen, 1818)
Chuvashia (Paramonov, 2016; Paramonov, 2017a).
Mordovia (Plavilshchikov, 1964; Antsiferova, Dobrosmyslov 1966; Ruchin & Pilipenko, 2015; Pilipenko et al., 2020).
Nephrotoma tenuipes (Riedel, 1910)
Chuvashia (Paramonov, 2018a). Mordovia (Pilipenko et al., 2020).
Nigrotipula nigra (Linnaeus, 1758)
Chuvashia (Paramonov, 2016; Paramonov, 2017a). Mordovia (Pilipenko et al., 2020). Saratov (Paramonov, 2015).
Prionocera pubescens Loew, 1844
Ulyanovsk (Paramonov, 2006).
Prionocera subserricornis (Zetterstedt, 1851)
Mordovia (Ruchin & Pilipenko, 2015; Pilipenko et al., 2020).
Prionocera turcica (Fabricius, 1787)
Mordovia (Ruchin & Pilipenko, 2015). Voronezh (Negrobov, 2005).
Tanyptera atrata (Linnaeus, 1758)
Chuvashia (Paramonov, 2016; Paramonov, 2017a). Mordovia (Ruchin & Pilipenko, 2015; Pilipenko et al., 2020). Samara (Rosenberg, 2007). Saratov (Paramonov, 2018b). Ulyanovsk (Paramonov, 2006).
Tanyptera nigricornis (Meigen, 1818)
Ulyanovsk (Paramonov, 2006).
Tipula fulvipennis De Geer, 1776
Chuvashia (Paramonov, 2018a). Mordovia (Ruchin & Pilipenko, 2015). Saratov (Paramonov, 2015). Ulyanovsk (Paramonov, 2006). Tipula luna Westhoff, 1879 Chuvashia (Paramonov, 2018a). Mordovia (Pilipenko et al., 2020). Saratov (Paramonov, 2015).
Tipula maxima Poda, 1761
Chuvashia (Paramonov, 2018a).
Saratov (Paramonov, 2019). Ulyanovsk (Paramonov, 2006).
Tipula unca Wiedemann, 1817
Chuvashia (Paramonov, 2018a).
Tipula flavolineata Meigen, 1804
Chuvashia (Paramonov, 2017a).
Tipula affinis Schummel, 1833
Mordovia (Pilipenko et al., 2020). Ulyanovsk (Paramonov, 2006).
Tipula fascipennis Meigen, 1818
Chuvashia (Paramonov, 2016; Paramonov, 2017a). Mordovia (Ruchin & Pilipenko, 2015; Pilipenko et al., 2020). Samara (Rosenberg, 2007).
Tipula laetabilis Zetterstedt, 1838
Saratov (Paramonov, 2015).
Tipula lunata Linnaeus, 1758
Chuvashia (Paramonov, 2018a).
Mordovia (Ruchin, 2008; Ruchin & Pilipenko, 2015; Pilipenko et al., 2020). Samara (Rosenberg, 2007). Saratov (Paramonov, 2015). Ulyanovsk (Paramonov, 2006).
Tipula mellea Schummel, 1833
Chuvashia (Paramonov, 2018a). Mordovia (Pilipenko et al., 2020). Saratov (Paramonov, 2015). Ulyanovsk (Paramonov, 2006).
Tipula peliostigma Schummel, 1833
Samara (Rosenberg, 2007). Saratov (Paramonov, 2018b). Tipula recticornis Schummel, 1833 Mordovia (Ruchin & Pilipenko, 2015).
Tipula selene Meigen, 1830
Saratov (Paramonov, 2015).
Tipula soosi Mannheims, 1954
Saratov (Paramonov, 2015, 2018b).
Tipula vernalis Meigen, 1804
Chuvashia (Paramonov, 2017a).
Mordovia (Plavilshchikov, 1964; Ruchin & Pilipenko, 2015; Pilipenko et al.,
2020).
Tipula nodicornis Meigen, 1818
Chuvashia (Paramonov, 2018a).
Tipula autumnalis Loew, 1864
Mordovia (Pilipenko et al., 2020).
Tipula luteipennis Meigen, 1830
Chuvashia (Paramonov, 2017a).
Mordovia (Ruchin & Pilipenko, 2015; Pilipenko et al., 2020).
Tipula jutlandica Nielsen, 1947
Mordovia (Pilipenko et al., 2020).
Tipula laetibasis Alexander, 1934
Mordovia (Pilipenko et al., 2020).
Tipula pseudovariipennis Czizek, 1912
Chuvashia (Paramonov, 2018a). Mordovia (Ruchin & Pilipenko, 2015).
Tipula submarmorata Schummel, 1833
Mordovia (Ruchin & Pilipenko, 2015; Pilipenko et al., 2020).
Tipula varipennis Meigen, 1818
Mordovia (Ruchin & Pilipenko, 2015; Pilipenko et al., 2020).
Tipula winthemi Lackschewitz, 1932
Mordovia (Pilipenko et al., 2020).
Tipula obsoleta Meigen, 1818
Chuvashia (Paramonov, 2018a).
Mordovia (Ruchin & Pilipenko, 2015; Pilipenko et al., 2020).
Tipula variicornis Schummel, 1833
Chuvashia (Paramonov, 2017a).
Tipula paludosa Meigen, 1830
Mordovia (Plavilshchikov, 1964; Ruchin, 2008; Ruchin & Pilipenko, 2015; Pilipenko et al., 2020).
Ulyanovsk (Paramonov, 2006). Voronezh (Negrobov, 2005).
Tipula subcunctans Alexander, 1921
Chuvashia (Paramonov, 2018a).
Mordovia (Pilipenko 2009; Ruchin & Pilipenko, 2015; Pilipenko et al., 2020). Saratov (Paramonov, 2018b).
Tipula hortorum Linnaeus, 1758
Chuvashia (Paramonov, 2018a).
Mordovia (Ruchin & Pilipenko, 2015; Pilipenko et al., 2020).
Tipula nubeculosa Meigen, 1804
Chuvashia (Paramonov, 2017a).
Mordovia (Ruchin & Pilipenko, 2015; Pilipenko et al., 2020). Ulyanovsk (Paramonov, 2006).
Tipula scripta Meigen, 1830
Chuvashia (Paramonov, 2016; Paramonov, 2017a). Mordovia (Ruchin & Pilipenko, 2015; Pilipenko et al., 2020). Samara (Rosenberg, 2007). Saratov (Paramonov, 2015). Ulyanovsk (Paramonov, 2006).
Tipula caesia Schummel, 1833
Saratov (Paramonov, 2015).
Tipula coerulescens Lackschewitz, 1923
Ulyanovsk (Paramonov, 2006).
Tipula couckei Tonnoir, 1921
Mordovia (Ruchin & Pilipenko, 2015). Saratov (Paramonov, 2015). Ulyanovsk (Paramonov, 2006).
Tipula lateralis Meigen, 1804
Chuvashia (Paramonov, 2018a).
Mordovia (Ruchin & Pilipenko, 2015; Pilipenko et al., 2020). Saratov (Paramonov, 2015). Ulyanovsk (Paramonov, 2006).
Tipula lucifera Savchenko, 1954
Chuvashia (Paramonov, 2017a).
Tipula pierrei Tonnoir, 1921
Mordovia (Pilipenko et al., 2020). Saratov (Paramonov, 2015). Ulyanovsk (Paramonov, 2006).
LIMONIIDAE
Achyrolimonia decemmaculata (Loew, 1873)
Chuvashia (Paramonov, 2018a). Mordovia (Esin et al., 2023).
Achyrolimonia neonebulosa (Alexander, 1924)
Mordovia (Pilipenko et al., 2020; Dvorak et al., 2020).
Arctoconopa melampodia (Loew, 1873)
Mordovia (Esin et al., 2023).
Atypophthalmus inustus (Meigen, 1818)
Mordovia (Esin et al., 2023). Saratov (Paramonov, 2015).
Austrolimnophila unica (Osten Sacken, 1869)
Mordovia (Esin et al., 2023).
Cheilotrichia imbuta (Meigen, 1818)
Mordovia (Esin et al., 2023).
Ryazan, Kasimov env., 54.94° N 41.34° E, N. Vikhrev, 17-19.06.2013, 1
Cheilotrichia cinerascens (Meigen, 1804)
Mordovia (Esin et al., 2023).
Cheilotrichia neglecta (Lackschewitz, 1927)
Mordovia (Pilipenko et al., 2020).
Chionea araneoides Dalman, 1816
Vladimir (Pavlov, 2017).
Chionea lutescens Lundstrom, 1907
Chuvashia (Paramonov, 2016, 2017b). Vladimir (Pavlov, 2006).
Dicranomyia autumnalis (Staeger, 1840)
Mordovia (Pilipenko et al., 2020).
Dicranomyia consimilis (Zetterstedt, 1838)
Mordovia (Pilipenko et al., 2020).
Dicranomyia danica Kuntze, 1919
Mordovia (Pilipenko et al., 2020).
Voronezh, Khoper SNR, Varvarino, 51.207° N, 41.719° E, on light, K. Tomkovich, 9-13.08.2022, 1 S.
Dicranomyia frontalis (Staeger, 1840)
Mordovia (Ruchin & Pilipenko, 2015; Pilipenko et al., 2020). Ryazan, Kasimov env., 54.94° N 41.34° E, N. Vikhrev, 17-19.06.2013, 1 21-26.07.2013, 1 S. Kasimov env., Oka River, 54.94° N, 41.34° E, N. Vikhrev, 21-24.07.2013, 2 S, 2 Tuma env., 55.143° N, 40.616° E, N. Vikhrev, 21.07.2013, 1
Dicranomyia halterata Osten Sacken, 1869
Saratov (Paramonov, 2018b).
Dicranomyia longipennis (Schummel, 1829)
Tambov, Tambov env., 52.677° N, 41.330° E, N. Vikhrev, 06.06.2012, 1
Dicranomyia modesta (Meigen, 1818)
Chuvashia (Paramonov, 2017a). Mordovia (Pilipenko et al., 2020).
Penza, Poim env., 53.0° N, 43.2° E, 27.07.2022, N. Vikhrev, 1S. Ryazan, Kasimov env., 54.94° N, 41.34° E, N. Vikhrev, 17-19.06.2013, 3 S. Kasimov env., Oka River, 54.94° N, 41.34° E, N. Vikhrev, 21-24.07.2013, 1 S. Tuma env., 55.143° N, 40.616° E, N. Vikhrev, 21.07.2013, 1 S.
Tambov, Tambov env., 52.677° N, 41.330° E, N. Vikhrev, 06.06.2012, 4 S. Volgograd, Mikhaylovka urban distr., Starosele env., 50.000° N, 43.235° E, N. Vikhrev, 07.06.2012, 1 S.
Dicranomyia morio (Fabricius, 1787)
Ryazan, Kasimov distr., Zalesnoe vill., 54.969° N, 41.327° E, N. Vikhrev, 2126.07.2013, 2 S.
Dicranomyia omissinervis de Meijere, 1918 Mordovia (Pilipenko et al., 2020).
Dicranophragma nemorale (Meigen, 1818)
Mordovia (Ruchin & Pilipenko, 2015; Pilipenko et al., 2020).
Dicranomyia schineriana (Alexander, 1964)
Chuvashia (Paramonov, 2017a). Mordovia (Pilipenko et al., 2020). Saratov (Paramonov, 2015).
Dicranophragma separatum (Walker, 1848)
Chuvashia (Paramonov, 2017a). Mordovia (Esin et al., 2023).
Dicranomyia tristis (Schummel, 1829)
Mordovia (Pilipenko et al., 2020). Saratov (Paramonov, 2018b).
Lipetsk, Stanovoe distr., Palenka vill., 52.887° N, 38.319° E, Quercetum, K. Tomkovich, 19.07.2014, 1
Dicranomyia ventralis (Schummel, 1829)
Mordovia (Pilipenko et al., 2020).
Dicranoptycha fuscescens (Schummel, 1829)
Chuvashia (Paramonov, 2017a). Mordovia (Pilipenko et al., 2020).
Ryazan, Kasimov distr., Zalesnoe vill., 54.969° N, 41.327° E, N. Vikhrev, 21-26.07.2013, 1$.
Saratov (Paramonov, 2015).
Discobola annulata (Linnaeus, 1758)
Saratov (Paramonov, 2018b).
Discobola caesarea (Osten Sacken, 1854)
Chuvashia (Paramonov, 2018a). Discobola parvispinula (Alexander, 1947)
Chuvashia (Paramonov, 2017a).
Mordovia (Pilipenko et al., 2020; Dvorak et al., 2020).
Elephantomyia krivosheinae Savchenko, 1976
Chuvashia (Paramonov, 2018a). Mordovia (Pilipenko et al., 2020).
Eloeophila maculata (Meigen, 1804)
Mordovia (Pilipenko et al., 2020).
Ryazan, Kasimov distr., Zalesnoe vill., 54.969° N, 41.327° E, N. Vikhrev, 21-26.07.2013, 1 S.
Epiphragma ocellare (Linnaeus, 1760)
Chuvashia (Paramonov, 2017a).
Mordovia (Ruchin & Pilipenko, 2015; Pilipenko et al., 2020). Samara (Rosenberg, 2007). Saratov (Paramonov, 2018b).
Erioconopa trivialis (Meigen, 1818)
Mordovia (Plavilshchikov, 1964; Ruchin & Pilipenko, 2015; Pilipenko et al., 2020). Tambov, Tambov env., 52.677° N, 41.330° E, N. Vikhrev, 06.06.2012, 4 S.
Erioptera distinguenda Stary, 1983
Tambov, Tambov env., 52.677° N, 41.330° E, N. Vikhrev, 06.06.2012, 1 S.
Erioptera flavata (Westhoff, 1882)
Mordovia (Esin et al., 2023).
Erioptera fuscipennis Meigen, 1818
Mordovia (Ruchin & Pilipenko, 2015).
Erioptera fusculenta Edwards, 1938
Tambov, Tambov env., 52.677° N, 41.330° E, N. Vikhrev, 6.06.2012, 4 S.
Erioptera griseipennis Meigen, 1838
Mordovia (Esin et al., 2023).
Erioptera lutea Meigen, 1804
Chuvashia (Paramonov, 2016). Mordovia (Ruchin & Pilipenko, 2015).
Volgograd, Serafimovich env., 49.57° N, 42.60° E, N. Vikhrev, 2829.04.2013, 1
Erioptera squalida Loew, 1871
Mordovia (Esin et al., 2023).
Erioptera bivittata (Loew, 1873)
Volgograd, Volgograd env., 48.572° N, 44.427° E, N. Vikhrev, 08.06.2012, 1
S, 1 ?.
Erioptera fossarum (Loew, 1873)
Volgograd, Novoanninskiy distr., Gulyaevskiy env., 50.418° N, 42.760° E, N. Vikhrev, 07.06.2012, 2 S.
Euphylidorea phaeostigma (Schummel, 1829)
Mordovia (Pilipenko et al., 2020).
Eutonia barbipes (Meigen, 1804)
Chuvashia (Paramonov, 2017a). Saratov (Paramonov, 2018b). Ulyanovsk (Paramonov, 2006).
Eutonia barbipes (Meigen, 1804)
Mordovia (Ruchin & Pilipenko, 2015; Pilipenko et al., 2020).
Gnophomyia viridipennis (Gimmerthal, 1847)
Mordovia (Esin et al., 2023).
Gonomyia abscondita Lackschewitz, 1935
Mordovia (Esin et al., 2023).
Gonomyia bifida Tonnoir, 1920
Tambov, Tambov env., 52.677° N, 41.330° E, Vikhrev, 06.06.2012, 2 S
Gonomyia lucidula de Meijere, 1920
Mordovia (Esin et al., 2023).
Gonomyia subtenella Savchenko, 1972
Mordovia, Mordovia SNR, Novenkiy cordon, 54.709° N, 43.214° E, on light, K. Tomkovich, 15-19.08.2022, 16-19.08.2022, 1 S, 3 S.
Ryazan, Kasimov distr., Zalesnoe vill., 54.969° N, 41.327° E, N. Vikhrev, 21-26.07.2013, 6 S, 2
Gonomyia abbreviata Loew, 1873
Mordovia (Esin et al., 2023).
Helius flavus (Walker, 1856)
Chuvashia (Paramonov, 2017a).
Tambov, Tambov env., 52.677° N, 41.330° E, N. Vikhrev, 06.06.2012, 1 S, 1 ?. Helius longirostris (Meigen, 1818)
Lipetsk, Lipetsk, 52.64° N, 39.70° E, Voronezh River, N. Vikhrev, 26.07.2022, 1 S. Mordovia (Pilipenko et al., 2020).
Ryazan, Kasimov distr., Zalesnoe vill., 54.969° N, 41.327° E, N. Vikhrev, 21-26.07.2013, 3 S, 2 Tuma env., 55.143° N, 40.616° E, N. Vikhrev, 21.07.2013, 4 S. Saratov (Paramonov, 2018b).
Hexatoma fuscipennis (Curtis, 1836)
Mordovia (Pilipenko et al., 2020).
Hexatoma nubeculosa (Burmeister, 1829)
Chuvashia (Paramonov, 2018a).
Hoplolabis vicina (Tonnoir, 1920)
Ryazan, Kasimov distr., Zalesnoe vill., 54.969° N, 41.327° E, N. Vikhrev, 21-26.07.2013, 2 S, 2
Hoplolabis yezoana (Alexander, 1924)
Chuvashia (Paramonov, 2017a).
Idioptera linnei Oosterbroek, 1992
Mordovia (Ruchin & Pilipenko, 2015; Pilipenko et al., 2020).
Idioptera pulchella (Meigen, 1830)
Mordovia (Ruchin & Pilipenko, 2015).
Ilisia maculata (Meigen, 1804)
Saratov (Paramonov, 2015).
Libnotes ladogensis (Lackschewitz, 1940)
Mordovia (Esin et al., 2023).
Limnophila pictipennis (Meigen, 1818)
Mordovia (Ruchin & Pilipenko, 2015).
Limnophila schranki Oosterbroek, 1992
Chuvashia (Paramonov, 2018a).
Mordovia (Ruchin & Pilipenko, 2015; Pilipenko et al., 2020). Ulyanovsk (Paramonov, 2006).
Volgograd, Serafimovich env., Don River, 49.6° N, 42.6° E, N. Vikhrev, 28.04.2013, 4 S. Serafimovich env., 49.57° N, 42.60° E, N. Vikhrev, 28-29.04.2013, 4 S.
Limonia albifrons (Meigen, 1818)
Mordovia (Ruchin & Pilipenko, 2015).
Limonia macrostigma (Schummel, 1829)
Mordovia (Pilipenko et al., 2020; Dvorak et al., 2020).
Ryazan, Kasimov distr., Zalesnoe vill., 54.969° N, 41.327° E, N. Vikhrev, 17-19.06.2013, 1 S. Kasimov env., 54.94° N, 41.34° E, N. Vikhrev, 21-26.07.2013, 1 S, 1 ?. Saratov (Paramonov, 2015).
Limonia nigropunctata (Schummel, 1829)
Chuvashia (Paramonov, 2016; Paramonov, 2017a). Mordovia (Pilipenko et al., 2020).
Limonia phragmitidis (Schrank, 1781)
Chuvashia (Paramonov, 2017a).
Mordovia (Ruchin & Pilipenko, 2015; Pilipenko et al., 2020). Ryazan, Kasimov distr., Zalesnoe vill., 54.969° N, 41.327° E, N. Vikhrev, 21-26.07.2013, 1 Kasimov env., 54.94° N, 41.34° E, N. Vikhrev, 17-19.06.2013, 1 S, 1 ?. Ranovo River, 53.72° N, 39.93° E, N. Vikhrev, 06.06.2012, 1 S. Saratov (Paramonov, 2015).
Limonia trivittata (Schummel, 1829)
Lipetsk, Stanovoe distr., Palenka vill., 52.887° N, 38.319° E, Quercetum, K. Tomkovich, 19.07.2014, 1
Mordovia (Ruchin & Pilipenko, 2015).
Metalimnobia bifasciata (Schrank, 1781)
Chuvashia (Paramonov, 2016; Paramonov, 2017a).
Mordovia (Ruchin & Pilipenko, 2015; Pilipenko et al., 2020; Dvorak et al.,
2020).
Nizhny Novgorod (Dvorak et al., 2022).
Ryazan, Tuma env., 55.143° N, 40.616° E, N. Vikhrev, 21.07.2013, 1 S.
Samara (Rosenberg, 2007).
Saratov (Paramonov, 2015).
Ulyanovsk (Paramonov, 2006; Dvorak et al., 2022).
Metalimnobia charlesi Salmela and Stary, 2009
Chuvashia (Paramonov, 2018a).
Metalimnobia quadrimaculata (Linnaeus, 1760)
Chuvashia (Paramonov, 2016; Paramonov, 2017a).
Mordovia (Ruchin & Pilipenko, 2015; Pilipenko et al., 2020; Dvorak et al., 2020). Nizhny Novgorod (Dvorak et al., 2022). Saratov (Paramonov, 2015; Dvorak et al., 2022). Ulyanovsk (Paramonov, 2006; Dvorak et al., 2022).
Metalimnobia quadrinotata (Meigen, 1818)
Chuvashia (Paramonov, 2016; Paramonov, 2017a). Mordovia (Ruchin & Pilipenko, 2015; Pilipenko et al., 2020). Ryazan, Ranovo River, 53.72° N, 39.93° E, N. Vikhrev, 06.06.2012, 1
Metalimnobia tenua Savchenko, 1976
Chuvashia (Paramonov, 2016; Paramonov, 2017a).
Molophilus appendiculatus (Staeger, 1840)
Mordovia (Esin et al., 2023).
Molophilus ater (Meigen, 1804)
Mordovia (Esin et al., 2023).
Molophilus bifidus Goetghebuer, 1920
Saratov (Paramonov, 2015).
Molophilus bihamatus de Meijere, 1918
Mordovia (Esin et al., 2023).
Molophilus crassipygus de Meijere, 1918
Mordovia (Esin et al., 2023). Saratov (Paramonov, 2015).
Molophilus griseus (Meigen, 1804)
Lipetsk, Lipetsk, 52.64° N, 39.70° E, Voronezh River, N. Vikhrev, 26.07.2022,
1 S.
Ryazan, Kasimov env., Syntul Lake, 54.94° N, 41.34° E, N. Vikhrev, 25.07.2013, 2 S.
Molophilus ochraceus (Meigen, 1818)
Mordovia (Ruchin & Pilipenko, 2015).
Molophilus propinquus (Egger, 1863)
Mordovia (Ruchin & Pilipenko, 2015).
Ryazan, Kasimov env., Syntul Lake, 54.94° N, 41.34° E, N. Vikhrev, 25.07.2013, 1 S.
Molophilus pullus Lackschewitz, 1927
Ulyanovsk, Radishchevo distr., Vyazovka env., 52.82° N, 48.35° E, 95-130 m a.s.l., Corydalis, K. Tomkovich, 2-5.05.2011, 2 S, 2
Volgograd, Serafimovich env., 49.57° N, 42.60° E, N. Vikhrev, 28-29.04.2013,
2 S, 2
Molophilus serpentiger Edwards, 1938
Mordovia (Esin et al., 2023).
Ormosia depilata Edwards, 1938
Mordovia (Esin et al., 2023). Ormosia lineata (Meigen, 1804)
Mordovia (Esin et al., 2023).
Phylidorea fulvonervosa (Schummel, 1829)
Chuvashia (Paramonov, 2018a). Mordovia (Pilipenko et al., 2020).
Phylidorea abdominalis (Staeger, 1840)
Mordovia (Esin et al., 2023).
Phylidorea bicolor (Meigen, 1804)
Mordovia (Pilipenko et al., 2020).
Phylidorea ferruginea (Meigen, 1818)
Chuvashia (Paramonov, 2018a).
Mordovia (Pilipenko et al., 2020).
Tambov, Tambov env., 52.677° N, 41.330° E, N. Vikhrev, 06.06.2012, 1 S.
Phylidorea longicornis (Schummel, 1829)
Mordovia (Esin et al., 2023).
Phylidorea squalens (Zetterstedt, 1838)
Mordovia (Pilipenko et al., 2020).
Pilaria nigropunctata (Agrell, 1945)
Mordovia (Esin et al., 2023).
Pilaria discicollis (Meigen, 1818)
Mordovia (Pilipenko et al., 2020).
Ryazan, Kasimov distr., Zalesnoe vill., 54.969° N, 41.327° E, N. Vikhrev, 21-26.07.2013, 1 S, 1 ?.
Pilaria fuscipennis (Meigen, 1818)
Mordovia (Pilipenko et al., 2020).
Pseudolimnophila lucorum (Meigen, 1818)
Chuvashia (Paramonov, 2017a). Mordovia (Esin et al., 2023). Saratov (Paramonov, 2015).
Tambov, Tambov env., 52.677° N, 41.330° E, N. Vikhrev, 6.06.2012, 1 S.
Pseudolimnophila sepium (Verrall, 1886)
Mordovia (Esin et al., 2023).
Volgograd, Volgograd env., 48.572° N, 44.427° E, N. Vikhrev, 8.06.2012, 1 S.
Rhabdomastix japonica Alexander, 1924
Mordovia (Esin et al., 2023).
Rhipidia maculata Meigen, 1818
Mordovia (Ruchin & Pilipenko, 2015; Pilipenko et al., 2020). Ulyanovsk (Paramonov, 2006).
Rhipidia uniseriata Schiner, 1864
Ulyanovsk (Dvorak et al., 2022). Mordovia (Esin et al., 2023).
Symplecta hybrida (Meigen, 1804)
Chuvashia (Paramonov, 2017a).
Mordovia (Ruchin & Pilipenko, 2015; Pilipenko et al., 2020).
Ryazan, Kasimov env., 54.94° N, 41.34° E, N. Vikhrev, 17-19.06.2013, 1 Kasimov env., Oka River, 54.94° N, 41.34° E, N. Vikhrev, 21-24.07.2013, 1 Saratov (Paramonov, 2015).
Tambov, Tambov env., 52.677° N, 41.330° E, N. Vikhrev, 06.06.2012, 1 S. Voronezh, Khoper SNR, Varvarino, 51.207° N, 41.719° E, on light, K. Tomkovich, 9-13.08.2022, 1 S. Ulyanovsk (Paramonov, 2006).
Volgograd, Volgograd env., 48.572° N, 44.427° E, N. Vikhrev, 08.06.2012,
1 S.
Symplecta pilipes (Fabricius, 1787)
Saratov (Paramonov, 2018b).
Tambov, Tambov env., 52.677° N, 41.330° E, N. Vikhrev, 06.06.2012, 1 S.
Tricyphona immaculata (Meigen, 1804)
Mordovia (Esin et al., 2023).
Tricyphona unicolor (Schummel, 1829)
Mordovia (Esin et al., 2023).
PEDICIIDAE
Dicranota gracilipes Wahlgren, 1905
Mordovia (Pilipenko et al., 2020).
Pedicia rivosa (Linnaeus, 1758)
Mordovia (Pilipenko et al., 2020). Ulyanovsk (Paramonov, 2006).
Tricyphona immaculata (Meigen, 1804)
Mordovia (Pilipenko et al., 2020).
Tricyphona unicolor (Schummel, 1829)
Mordovia (Pilipenko et al., 2020).
Ula bolitophila Loew, 1869
Mordovia (Pilipenko et al., 2020).
Ryazan, Kasimov distr., Zalesnoe vill., 54.969° N, 41.327° E, N. Vikhrev, 21-26.07.2013, 1$.
Ula sylvatica (Meigen, 1818)
Mordovia (Esin et al., 2023).
MYCETOPHILIDAE
Acnemia nitidicollis (Meigen, 1818)
Voronezh, Khoper River. x P22 road, 51.36° N, 42.04° E, N. Vikhrev, 27.04.2022, 2 S.
Allodia alternans (Zetterstedt, 1838)
Ulyanovsk, Radishchevo distr., Vjazovka env., 52.82° N, 48.35° E, 33-170 m a.s.l., K. Tomkovich, 1-9.05.2011, 1 S.
Allodia ornaticollis (Meigen, 1818)
Chuvashia (Borisova et al., 2018).
Allodia neglecta Edwards, 1925
Mordovia (Esin et al., 2023).
Allodiopsis domestica (Meigen, 1830)
Mordovia (Esin et al., 2023).
Allodia silvatica (Landrock, 1912)
Mordovia (Esin et al., 2023).
Anatella unguigera Edwards, 1921
Mordovia (Esin et al., 2023).
Boletina anderschi (Stannius, 1831)
Ulyanovsk, Radishchevo distr., Vjazovka env., 52.82° N, 48.35° E, 33-170 m a.s.l., K. Tomkovich, 2-8.05.2011, 5 S.
Boletina basalis (Meigen, 1818)
Mordovia (Esin et al., 2023).
Boletina cincticornis (Walker, 1848)
Mordovia (Esin et al., 2023).
Boletina dispecta Dziedzicki, 1885
Mordovia (Esin et al., 2023).
Boletina dubia (Meigen, 1804)
Mordovia (Esin et al., 2023).
Boletina gripha Dziedzicki, 1885
Mordovia (Esin et al., 2023).
Ulyanovsk, Radishchevo distr., Vjazovka env., 52.82° N, 48.35° E, 33-170 m a.s.l., K. Tomkovich, 02-08.05.2011, 1 S.
Voronezh, Khoper River. x P22 road, 51.36° N, 42.04° E, N. Vikhrev, 27.04.2022, 16
Boletina moravica Landrock, 1912
Mordovia (Esin et al., 2023).
Boletina nigricoxa Staeger, 1840
Mordovia (Esin et al., 2023).
Boletina trivittata (Meigen, 1818)
Mordovia (Esin et al., 2023).
Brevicornu fuscipenne (Stager, 1840)
Mordovia (Esin et al., 2023).
Brevicornu sericoma (Meigen, 1830)
Mordovia (Esin et al., 2023).
Cordyla brevicornis (Stager, 1840)
Mordovia (Esin et al., 2023).
Cordyla crassicornis Meigen, 1818
Mordovia (Esin et al., 2023).
Cordyla fasciata Meigen, 1830
Mordovia (Esin et al., 2023).
Cordyla flaviceps (Stager, 1840)
Mordovia (Esin et al., 2023).
Cordyla fusca Meigen, 1804
Mordovia (Esin et al., 2023).
Cordyla insons Lastovka & Matile, 1974
Mordovia (Esin et al., 2023).
Cordyla murina Winnertz, 1863
Mordovia (Esin et al., 2023).
Cordyla nitidula Edwards, 1925
Mordovia (Esin et al., 2023).
Cordyla sixi (Barendrecht, 1938)
Mordovia (Esin et al., 2023).
Dynatosoma thoracicum (Zetterstedt, 1838)
Mordovia (Esin et al., 2023).
Epicypta aterrima (Zetterstedt, 1852)
Mordovia (Esin et al., 2023).
Exechia dorsalis (Stager, 1840)
Mordovia (Esin et al., 2023).
Epicypta fumigata (Dziedzicki, 1923)
Mordovia (Esin et al., 2023).
Exechia fusca (Meigen, 1804)
Mordovia (Esin et al., 2023).
Exechia nigroscutellata Landrock, 1912
Chuvashia (Borisova et al., 2018).
Exechia repanda Johannsen, 1912
Mordovia (Esin et al., 2023).
Exechia separata Lundstrom, 1912
Mordovia (Esin et al., 2023).
Greenomyia mongolica Lastovka & Matile, 1974
Mordovia (Esin et al., 2023).
Leia bimaculata (Meigen, 1804)
Voronezh, Khoper River. x P22 road, 51.36° N, 42.04° E, N. Vikhrev, 27.04.2022, 1 S.
Leia fascipennis Meigen, 1818
Mordovia (Esin et al., 2023).
Mycetophila alea Laffoon, 1965
Mordovia (Esin et al., 2023).
Mycetophila attonsa Laffoon, 1957
Mordovia (Esin et al., 2023).
Mycetophila fungorum (De Geer, 1776)
Chuvashia (Borisova et al., 2018).
Mordovia (Plavilshchikov, 1964, as Fungivora fungorum).
Mycetophila luctuosa Meigen, 1830
Chuvashia (Borisova et al., 2018).
Mycetophila marginata Winnertz, 1863
Chuvashia (Borisova et al., 2018).
Mycetophila sepulta (Laffoon, 1957)
Mordovia (Esin et al., 2023).
Mycetophila sigillata Dziedzicki, 1884
Voronezh, Khoper River. x P22 road, 51.36° N, 42.04° E, N. Vikhrev, 27.04.2022, 1 S.
Mycetophila signata Meigen, 1830
Mordovia (Esin et al., 2023).
Mycetophila stolida Walker, 1856
Mordovia (Esin et al., 2023).
Mycetophila strigatoides Landrock, 1927
Mordovia (Esin et al., 2023).
Mycetophila triangulata Dziedzicki, 1884
Mordovia (Esin et al., 2023).
Mycetophila unipunctata Meigen, 1818
Mordovia (Esin et al., 2023).
Mycomya annulata (Meigen, 1818)
Mordovia (Esin et al., 2023).
Mycomya occultans (Winnertz, 1863)
Mordovia (Esin et al., 2023).
Mycomya tridens (Lundstrom, 1911)
Ulyanovsk, Radishchevo distr., Vjazovka env., 52.82° N, 48.35° E, 33-170 m a.s.l., K. Tomkovich, 2-8.05.2011, 1 S.
Neoempheria brevilineata Okada, 1939
Mordovia (Esin et al., 2023).
Neoempheria pictipennis (Haliday, 1833)
Mordovia (Esin et al., 2023).
Platurocypta testata (Edwards, 1925)
Mordovia (Esin et al., 2023).
Rondaniella dimidiata (Meigen, 1804)
Mordovia (Esin et al., 2023).
Rymosia signatipes (van der Wulp, 1859)
Mordovia (Esin et al., 2023).
Sceptonia costata (van der Wulp, 1859)
Mordovia (Esin et al., 2023).
Sceptonia demeijerei Bechev, 1997
Mordovia (Esin et al., 2023).
Sceptonia fumipes Edwards, 1925
Mordovia (Esin et al., 2023).
Sceptonia nigra (Meigen, 1804)
Mordovia (Esin et al., 2023).
Synplasta dulcia (Dziedzicki, 1910)
Mordovia (Esin et al., 2023).
Synplasta exclusa (Dziedzicki, 1910)
Mordovia (Esin et al., 2023).
Synplasta rufilatera (Edwards, 1941)
Ulyanovsk, Radishchevo distr., Vjazovka env., 52.82° N, 48.35° E, 33-170 m a.s.l., K. Tomkovich, 1-9.05.2011, 1 S.
Trichonta atricauda (Zetterstedt, 1852)
Voronezh, Khoper River. x P22 road, 51.36° N, 42.04° E, N. Vikhrev, 27.04.2022, 1 S.
Zygomyia notata (Stannius, 1831)
Mordovia (Esin et al., 2023).
Zygomyia pictipennis (Stager, 1840)
Mordovia (Esin et al., 2023).
Zygomyia valida Winnertz, 1863
Mordovia (Esin et al., 2023).
KEROPLATIDAE
Isoneuromyia semirufa (Meigen, 1818)
Mordovia (Esin et al., 2023).
Keroplatus testaceus Dalman, 1818
Mordovia (Esin et al., 2023).
Keroplatus tipuloides Bosc, 1792
Mordovia (Esin et al., 2023).
Macrocera centralis Meigen, 1818
Mordovia (Esin et al., 2023).
Monocentrota lundstromi Edwards, 1925
Mordovia (Esin et al., 2023).
Neoplatyura flava (Macquart, 1826)
Mordovia (Esin et al., 2023).
Neoplatyura modesta (Winnertz, 1863)
Mordovia (Esin et al., 2023).
Orfelia fasciata (Meigen, 1804)
Volgograd, Volgograd env., 48.572° N, 44.427° E, saltish pond edge, K. Tomkovich, 08.06.2012, 1 1
Orfelia lugubris (Zetterstedt, 1851)
Mordovia (Esin et al., 2023).
Pyratula zonata (Zetterstedt, 1855)
Mordovia (Esin et al., 2023).
PTYCHOPTERIDAE
Ptychoptera albimana Fabricius, 1781
Mordovia (Esin et al., 2023). Saratov (Paramonov, 2004)
Ptychoptera contaminata (Linnaeus, 1758)
Lipetsk, Voronezh River, 52.64° N, 39.7° E, N. Vikhrev, 26.07.2022, 1 Mordovia (Feoktistov, 2011). Saratov (Paramonov, 2004).
Voronezh, Khoper SNR, Varvarino, 51.207° N, 41.719° E, floodplain, K. Tomkovich, 9-13.08.2022, 1 S, 1 ?.
Ptychoptera lacustris Meigen, 1830
Saratov (Paramonov, 2004). Tatarstan (Paramonov, 2010). Volgograd (Paramonov, 2004).
Ptychoptera scutellaris Meigen, 1818
Tatarstan (Paramonov, 2014).
SUBORDER BRACHYCERA MUSCIDAE
Achanthiptera rohrelliformis Robineau-Desvoidy, 1830
Mordovia (Vikhrev et al., 2020). Samara (Lyubvina, 2016a).
Azelia aterrima Meigen, 1826
Mordovia (Vikhrev et al., 2020).
Voronezh, Khoper SNR, Varvarino, 51.21° N, 41.72° E, K. Tomkovich, 4-5.08.2022, 1 S.
Azelia cilipes Haliday, 1838
Mordovia (Vikhrev et al., 2020). Penza (Lyubvina, 2016b). Tatarstan (Yuzekaeva, 2019).
Azelia gibbera Meigen, 1826
Mordovia (Vikhrev et al., 2020).
Azelia monodactyla Loew, 1874
Mordovia (Vikhrev et al., 2020).
Azelia nebulosa Robineau-Desvoidy, 1830
Mordovia (Vikhrev et al., 2020).
Azelia spinosa Vikhrev, 2015
Volgograd (Vikhrev, 2015a).
Azelia trigonica Hennig, 1956
Mordovia (Vikhrev et al., 2020).
Azelia zetterstedtii Rondani, 1866
Mordovia (Vikhrev et al., 2020). Tatarstan (Yuzekaeva, 2019).
Coenosia agromyzina Fallen, 1825
Mordovia (Vikhrev et al., 2020).
Ryazan, Kasimov env., 54.94° N, 41.34° E, N. Vikhrev, 17-19.07.2013, 3S, 1 ?. Samara (Lyubvina, 2016a).
Voronezh, Khoper SNR, Varvarino, 51.21° N, 41.72° E, K. Tomkovich, 13.08.2022, 1 S.
Coenosia ambulans Meigen, 1826
Samara (Lyubvina, 2016a).
Coenosia atra Meigen, 1830
Mordovia (Vikhrev et al., 2020).
Coenosia humilis Meigen, 1826
Mordovia (Vikhrev et al., 2020).
Coenosia lineatipes Zetterstedt, 1845
Mordovia, Krasnoslobodsk distr., Selischi, 54.481° N, 43.522° E, forest edge, YPT, 1-4.08.2020, K. Tomkovich, M. Esin, 1 S, identified by V. Sorokina.
Coenosia mollicula Fallen, 1825
Mordovia (Vikhrev et al., 2020).
Nizhny Novgorod, W of Nizhny Novgorod (56.28° N, 43.70° E), birch-pin forest, 12.07.2009, N. Vikhrev, 2$.
Ryazan, Tuma env., 55.143° N, 40.616° E, N. Vikhrev, 21.07.2013, 2 S. Samara (Lyubvina, 2016a).
Ulyanovsk, Staroe Eryomkino (54.32° N, 49.28° E), V.B. Isaeva, 24.07.1998, 1 Voronezh, Khoper SNR, Varvarino, 51.21° N, 41.72° E, K. Tomkovich, 4-5.08.2022, 1 S.
Coenosia paludis Tiensuu, 1939
Samara (Lyubvina, 2016a).
Coenosia pudorosa Collin, 1953
Mordovia (Vikhrev et al., 2020).
Coenosia pumila Fallen, 1825
Mordovia (Vikhrev et al., 2020).
Nizhny Novgorod, Dzerzhinsk env., 56.21° N, 43.62° E, 15.08.2009, N. Vikhrev, 4 S.
Penza, Poim env., 53.0° N, 43.2° E, 27.07.2022, N. Vikhrev, 2 S. Samara (Lyubvina, 2016a).
Voronezh, Khoper SNR, Varvarino, 51.21° N, 41.72° E, K. Tomkovich, 13.08.2022, 1 S.
Coenosia pygmaea Zetterstedt, 1845
Mordovia (Vikhrev et al., 2020).
Coenosia rufipalpis Meigen, 1826
Mordovia (Vikhrev et al., 2020). Samara (Lyubvina, 2016a).
Coenosia strigipes Stein, 1916
Mordovia (Vikhrev et al., 2020). Penza (Lyubvina, 2016b).
Voronezh, Khoper SNR, Varvarino, 51.21° N, 41.72° E, K. Tomkovich, 13.08.2022, 1 S.
Coenosia testacea Robineau-Desvoidy, 1830
Mordovia (Vikhrev et al., 2020). Samara (Lyubvina, 2016a).
Tambov, W of Kirsanov, 52.65° N, 42.62° E, 27-28.07.2022, N. Vikhrev, 1 S. Coenosia tigrina Fabricius, 1775
Volgograd, saltish pond, 48.572° N, 44.427° E, 8.06.2012, N. Vikhrev, 1 S. Voronezh, Khoper SNR, Varvarino, 51.21° N, 41.72° E, K. Tomkovich, 4-5.08.2022, 2
Coenosia trilineela Zetterstedt, 1838
Mordovia (Vikhrev et al., 2020). Coenosia verralli Collin, 1953
Mordovia (Vikhrev et al., 2020).
Nizhny Novgorod, Dzerzhinsk env., 56.21° N, 43.62° E, 16-18.08.2009, N. Vikhrev, 4 S.
Samara (Lyubvina, 2016a).
Drymeia vicana Harris, 1780
Mordovia (Vikhrev et al., 2020). Samara (Rosenberg, 2007). Voronezh (Negrobov, 2005).
Eginia ocypterata Meigen, 1826
Mordovia (Vikhrev et al., 2020).
Nizhny Novgorod, W of Nizhny Novgorod, 56.28° N43.70° E, birch-pin forest, 12.07.2009, N. Vikhrev, 4 S.
Samara (Rosenberg, 2007).
Eudasyphora cyanicolor Zetterstedt, 1845
Mordovia (Vikhrev et al., 2020). Samara (Lyubvina, 2016a).
Voronezh, Khoper SNR, Dubovaya Khata, 51.25° N, 41.78° E, K. Tomkovich, 6.08.2022, 2
Graphomya maculata Scopoli, 1763
Mordovia (Vikhrev et al., 2020). Nizhny Novgorod (Vikhrev, 2011a). Penza (Lyubvina, 2016b). Samara (Vikhrev, 2011a).
Tambov, W of Kirsanov, 52.65° N, 42.62° E, 27-28.07.2022, N. Vikhrev, 1 Ulyanovsk (Vikhrev, 2011a). Voronezh (Negrobov, 2005).
Gymnodia humilis Zetterstedt, 1860
Mordovia (Vikhrev et al., 2020).
Gymnodia polystigma Meigen, 1826
Mordovia (Vikhrev et al., 2020).
Haematobia irritans Linnaeus, 1758
Mordovia (Vikhrev et al., 2020). Samara (Lyubvina, 2016a). Tambov (Molev et al., 1967). Voronezh (Negrobov, 2005).
Haematobosca stimulans Meigen, 1824
Mordovia (Vikhrev et al., 2020). Samara (Lyubvina, 2016a). Voronezh (Negrobov, 2005).
Haematobia titillans Bezzi, 1907
Tambov, Borschevka vill. (52.6° N, 41.3° E), pasture, calf, 27.08.1964, G. Veselkin, 1 S, 2
Hebecnema umbratica Meigen, 1826
Mordovia (Vikhrev et al., 2020).
Hebecnema vespertina Fallen, 1823
Mordovia (Vikhrev et al., 2020).
Helina allotalla (Meigen, 1830)
Samara (Lyubvina, 2016a).
Helina cilipes (Schnabl, 1889)
Penza (Lyubvina, 2016b).
Helina ciliatocosta (Zetterstedt, 1845)
Penza (Lyubvina, 2016b). Samara (Rosenberg, 2007).
Helina confinis Fallen, 1825
Mordovia (Vikhrev et al., 2020).
Lipetsk, Lipetsk, 52.64° N, 39.70° E, Voronezh River, N. Vikhrev, 26.07.2022, 1 Penza (Lyubvina, 2016b). Samara (Lyubvina, 2016a).
Voronezh, Khoper SNR, Varvarino, 51.21° N, 41.72° E, K. Tomkovich, 4-5.08.2022, 2
Helina cothurnata Rondani, 1866
Mordovia (Vikhrev et al., 2020).
Helina evecta Harris, 1780
Mordovia (Vikhrev et al., 2020).
Samara, Enduraykino vill., 53.8° N 51.38° E, 12.08.22, A. Kuzovenko, 2 2 Voronezh, Khoper SNR, Varvarino, 51.21° N, 41.72° E, K. Tomkovich, 45.08.2022, 1 1
Helina depuncta Fallen, 1825
Mordovia (Vikhrev et al., 2020).
Lipetsk, Lipetsk, 52.64° N, 39.70° E, Voronezh River, N. Vikhrev, 26.07.2022, 1 Samara (Rosenberg, 2007).
Helina impuncta Fallen, 1825
Mordovia (Vikhrev et al., 2020). Penza (Lyubvina, 2016b). Samara (Rosenberg, 2007).
Voronezh, Khoper SNR, Dubovaya Khata, 51.25° N, 41.78° E, K. Tomkovich, 6.08.2022, 1
Helina laxifrons (Zetterstedt, 1860)
Penza (Lyubvina, 2016b).
Samara (Lyubvina, 2016a). Vladimir (Pavlov, 2006).
Helina maculipennis Zetterstedt, 1845
Mordovia (Vikhrev et al., 2020).
Helina moedlingensis (Schnabl 1911)
Penza (Lyubvina, 2016b). Samara (Lyubvina, 2016a).
Helina obscurata Meigen, 1826
Mordovia (Vikhrev et al., 2020). Penza (Lyubvina, 2016b). Samara (Lyubvina, 2016a).
Helina parcepilosa (Stein, 1907)
Penza (Lyubvina, 2016b). Samara (Lyubvina, 2016a).
Helina pertusa Meigen, 1826
Mordovia (Vikhrev et al., 2020).
Helina pubescens (Stein, 1893)
Penza (Lyubvina, 2016b).
Helina quadrum (Fabricius, 1805)
Samara (Lyubvina, 2016a).
Helina quadrinotata (Meigen, 1826)
Penza (Lyubvina, 2016b).
Helina reversio (Harris, 1780)
Samara (Lyubvina, 2016a).
Helina setiventris Ringdahl, 1924
Mordovia (Vikhrev et al., 2020). Samara (Lyubvina, 2016a).
Helina sexmaculata Preyssler, 1791
Mordovia (Vikhrev et al., 2020).
Samara, Enduraykino vill., 53.8° N 51.38° E, 12.08.2022, A. Kuzovenko, 1 S. Voronezh, Khoper SNR, Varvarino, 51.21° N, 41.72° E, K. Tomkovich, 9-13.08.2022, 1 S, 1 ?.
Helina spinicosta (Zetterstedt, 1845)
Samara (Lyubvina, 2016a).
Helina tetrastigma Meigen, 1826
Mordovia (Vikhrev et al., 2020). Penza (Lyubvina, 2016b).
Helina trivittata Zetterstedt, 1860
Mordovia (Vikhrev et al., 2020).
Samara, Enduraykino vill., 53.8° N 51.38° E, 12.08.22, A. Kuzovenko, 1 Hydrotaea aenescens Wiedemann, 1830
Chuvashia, Yadrin env., 55.88° N, 46.29° E, N. Vikhrev, 15.07.2009, 1 S. Mordovia (Vikhrev et al., 2020).
Ulyanovsk, Shilovka vill. env., 52.82° N, 48.35° E, 1-3.09.2010, K. Tomkovich,
3S, 1 ?.
Hydrotaea albipuncta Zetterstedt, 1845
Chuvashia, Yadrin env., 55.88° N, 46.29° E, N. Vikhrev, 13.07.2009, 2 S. Samara (Lyubvina, 2016a).
Hydrotaea armipes Fallen, 1825
Mordovia (Vikhrev et al., 2020). Penza (Lyubvina, 2016b).
Hydrotaea basdeni Collin, 1939
Chuvashia, Yadrin env., 55.88° N, 46.29° E, N. Vikhrev, 13.07.2009, 1 S, 1 ?.
Hydrotaea borussica Stein, 1899
Mordovia (Vikhrev et al., 2020).
Hydrotaea capensis Wiedemann, 1818
Mordovia (Vikhrev et al., 2020).
Hydrotaea cyrtoneurina Zetterstedt, 1845
Mordovia (Vikhrev et al., 2020).
Vladimir, S of Vladimir, 56.06° N, 40.46° E, 5.06.2011, N. Vikhrev, 1 S. Voronezh, Khoper SNR, Varvarino, 51.21° N, 41.72° E, K. Tomkovich, 9-13.08.2022, 2 S.
Hydrotaea dentipes Fabricius, 1805
Mordovia (Vikhrev et al., 2020).
Lipetsk, Lipetsk, 52.64° N, 39.70° E, Voronezh River, N. Vikhrev, 26.07.2022, 1 S. Penza (Levkovich & Levkovich, 1979).
Samara (Lyubvina, 2016a).
Tambov, W of Kirsanov, 52.65° N, 42.62° E, 27-28.07.2022, N. Vikhrev, 1 S.
Hydrotaea diabolus Harris, 1780
Mordovia (Vikhrev et al., 2020).
Hydrotaea floccosa Macquart, 1835
Chuvashia, Yadrin env., 55.88° N, 46.29° E, N. Vikhrev, 13.07.2009, 2 S. Mordovia (Vikhrev et al., 2020).
Hydrotaea glabricula Fallen, 1825
Mordovia (Vikhrev et al., 2020).
Hydrotaea ignava Harris, 1780
Chuvashia, Yadrin env., 55.88° N, 46.29° E, N. Vikhrev, 13.07.2009, 1 Mordovia (Vikhrev et al., 2020). Penza (Levkovich & Levkovich, 1979).
Ryazan, Kasimov env., 54.94° N, 41.34° E, N. Vikhrev, 17-19.07.2013, 1 Samara (Rosenberg, 2007).
Voronezh, Khoper SNR, Dubovaya Khata, 51.25° N, 41.78° E, K. Tomkovich, 6-8.08.2022, 1
Hydrotaea irritans Fallen, 1823
Mordovia (Vikhrev et al., 2020). Ryazan (Vikhrev, 2013). Samara (Lyubvina, 2016a).
Hydrotaea meteorica Linnaeus, 1758
Mordovia (Vikhrev et al., 2020). Samara (Lyubvina, 2016a).
Hydrotaea militaris Meigen, 1826
Mordovia (Vikhrev et al., 2020).
Hydrotaea nidicola Malloch, 1925
Voronezh (Tewelde, Gaponov, 2019).
Hydrotaea palaestrica Meigen, 1826
Mordovia (Vikhrev et al., 2020). Penza (Lyubvina, 2016b).
Hydrotaea pandellei Stein, 1899
Mordovia (Vikhrev et al., 2020).
Hydrotaea parva Meade, 1889
Mordovia (Vikhrev et al., 2020).
Hydrotaea pellucens Porchinskiy, 1879
Mordovia (Vikhrev et al., 2020). Voronezh (Negrobov, 2005).
Hydrotaea pilipes Stein, 1903
Chuvashia, Yadrin env., 55.88° N, 46.29° E, N. Vikhrev, 15.07.2009, 1 Mordovia (Vikhrev et al., 2020).
Ryazan, Kasimov env., Zelenoe, 54.94° N, 41.34° E, N. Vikhrev, 17-19.07.2013, 1
Hydrotaea similis Meade, 1887
Mordovia (Vikhrev et al., 2020).
Hydrotaea tuberculata Rondani, 1866
Mordovia (Vikhrev et al., 2020).
Vladimir reg., S of Vladimir, 56.06° N, 40.46° E, 5.06.2011, N. Vikhrev, 1 S.
Hydrotaea unispinosa Stein, 1898
Mordovia (Esin et al., 2023).
Hydrotaea velutina Robineau-Desvoidy, 1830
Mordovia (Vikhrev et al., 2020).
Tambov, W of Tambov, 52.677° N, 41.330° E, N. Vikhrev, 6.06.2012, 1 S, 1 ?.
Limnophora maculosa Meigen, 1826
Mordovia (Vikhrev et al., 2020). Samara (Lyubvina, 2016a). Voronezh (Negrobov, 2005).
Limnophora nigripes Robineau-Desvoidy, 1830
Mordovia (Esin et al., 2023).
Limnophora pollinifrons Stein, 1916
Chuvashia, Yadrin env., 55.925° N, 46.288° E, Sura River, N. Vikhrev, 15.07.2009, 5 S, 3
Mordovia (Vikhrev et al., 2020).
Nizhny Novgorod, Dzerzhinsk env., 56.21° N, 43.62° E, 18.08.2009, N. Vikhrev, 2 S.
Saratov, Saratov, Zakhvatkin, 27.07.1931, 1 S.
Ulyanovsk, Shilovka vill. env., 52.82° N, 48.35° E, 1-3.09.2010, K. Tomkovich, 4 S, 4
Limnophora riparia Fallen, 1824
Chuvashia, Yadrin env., 55.88° N, 46.29° E, N. Vikhrev, 15.07.2009, 4 S, 1 ?. Mordovia (Vikhrev et al., 2020).
Vladimir, S of Vladimir, 56.055 ° N, 40.404° E, 5.06.2011, N. Vikhrev, 3 S, 3 Limnophora rufimana Strobl, 1893
Volgograd, (Medveditsa R.) 50.00° N, 43.235° E, N. Vikhrev, 7.06.2012, 2 S.
Limnophora tigrina Am Stein, 1860
Mordovia (Vikhrev et al., 2020).
Nizhny Novgorod, Dzerzhinsk env., 56.21° N, 43.62° E, 18-19.08.2009, N. Vikhrev, 4 S, 2
Ulyanovsk, Shilovka vill. env., 52.82° N, 48.35° E, 1-3.09.2010, K. Tomkovich,
1 S, 1 ?.
Voronezh, 20 km W of Liski (51.00° N, 39.2° E), K. Tomkovich, 8-9.07.2012,
1 ?.
Limnophora triangula Fallen, 1825
Mordovia (Vikhrev et al., 2020).
Nizhny Novgorod, Dzerzhinsk env., 56.21° N, 43.62° E, 18.08.2009, N. Vikhrev, 1
Lispe brunnicosa Becker, 1904
Volgograd (Vikhrev 2012a).
Lispe caesia Meigen, 1826
Volgograd (Vikhrev, 2016).
Lispe consanguinea Loew, 1858
Chuvashia (Vikhrev, 2011b). Mordovia (Vikhrev et al., 2020). Nizhnyi Novgorod (Vikhrev, 2011b). Ryazan (Vikhrev, 2014). Samara (Rosenberg, 2007). Saratov (Vikhrev, 2014). Ulyanovsk (Vikhrev, 2011b). Vladimir (Vikhrev, 2014). Volgograd (Vikhrev, 2014). Voronezh (Vikhrev, 2014).
Lispe hebeiensis Ma & Tian, 1993
Volgograd (Vikhrev, 2015b).
Lispe loewi Ringdahl, 1922
Volgograd (Vikhrev, 2015b).
Lispe longicollis Meigen 1826
Saratov (Vikhrev, 2012b). Volgograd (Vikhrev, 2012b).
Lispe melaleuca Loew, 1847
Mordovia (Vikhrev et al., 2020). Ryazan (Vikhrev, 2015b). Saratov (Vikhrev, 2015b). Volgograd (Vikhrev, 2015b).
Lispe nana Macquart, 1835
Chuvashia (Vikhrev, 2014). Mordovia (Vikhrev et al., 2020). Ryazan (Vikhrev, 2014).
Lispe neimongola Tian & Ma, 2000
Volgograd (Vikhrev, 2015b).
Lispe nubilipennis Loew, 1873
Volgograd (Vikhrev, 2014).
Lispe odessae Becker, 1904
Volgograd (Vikhrev, 2016).
Lispe pygmaea Fallen, 1825
Mordovia (Vikhrev et al., 2020). Nizhny Novgorod (Vikhrev, 2016). Samara (Rosenberg, 2007). Volgograd (Vikhrev, 2016). Voronezh (Vikhrev, 2016). Ulyanovsk (Vikhrev, 2016).
Lispe parcespinosa bohemica Becker, 1904
Chuvashia (Vikhrev, 2015b). Nizhnyi Novgorod (Vikhrev, 2015b). Ryazan (Vikhrev, 2015b).
Lispe uliginosa (Fallén, 1825)
Samara (Rosenberg, 2007).
Lispe superciliosa Loew, 1861
Mordovia (Vikhrev et al., 2020). Saratov (Vikhrev, 2015b). Tambov (Vikhrev, 2015b). Ulyanovsk (Vikhrev, 2015b). Volgograd (Vikhrev, 2015b).
Lispe tentaculata De Geer, 1776
Chuvashia (Vikhrev, 2011b). Mordovia (Vikhrev et al., 2020). Nizhnyi Novgorod (Vikhrev, 2011b). Vladimir (Vikhrev, 2011b). Voronezh (Negrobov, 2005). Ulyanovsk (Vikhrev, 2011b).
Lispocephala alma Meigen, 1826
Mordovia (Vikhrev et al., 2020). Vladimir (Pavlov, 2006).
Lispocephala erythrocera Robineau-Desvoidy, 1830
Mordovia (Vikhrev et al., 2020).
Lipetsk, Lipetsk, 52.64° N, 39.70° E, Voronezh River, N. Vikhrev, 26.07.2022, 1 S. Ryazan reg., Ranova River, 53.72° N, 39.93° E, N. Vikhrev, 6.06.2012, 5 S,
2 ?.
Samara (Lyubvina, 2016a).
Tambov, W of Kirsanov, 52.65° N, 42.62° E, 27-28.7.2022, N. Vikhrev, 1 S. Ulyanovsk, Ulyanovsk, Sviyaga River, 54.33° N, 48.34° E, 25.04.2011, K. Tomkovich, 1 S, 1 ?.
Vladimir (Pavlov, 2006).
Volgograd reg., Kalach-na-Donu env., 48.65° N, 43.60° E, 30.04.2013, N. Vikhrev, 1 S, 2
Lispocephala verna Fabricius, 1794
Mordovia (Vikhrev et al., 2020).
Macrorchis meditata Fallen, 1825
Vladimir, S of Vladimir, 56.055° N, 40.404° E, 15.08.2009, N. Vikhrev, 2 S.
Mesembrina meridiana Linnaeus, 1758
Mordovia (Vikhrev et al., 2020). Penza (Lyubvina, 2016b). Samara (Lyubvina, 2016a). Voronezh (Negrobov, 2005).
Mesembrina mystacea Linnaeus, 1758
Mordovia (Vikhrev et al., 2020). Samara (Lyubvina, 2016a).
Morellia aenescens Robineau-Desvoidy, 1830
Mordovia (Vikhrev et al., 2020). Voronezh (Negrobov, 2005).
Morellia hortorum Fallen, 1817
Chuvashia, Yadrin env., 55.88° N, 46.29° E, N. Vikhrev, 13.07.2009, 1 S. Penza (Lyubvina, 2016b). Samara (Rosenberg, 2007).
Ulyanovsk, Sura distr., Nikitio, 54.46° N, 47.07° E, 19.07.2001, 1 S. Voronezh (Negrobov, 2005).
Morellia podagrica Loew, 1857
Mordovia (Vikhrev et al., 2020). Samara (Rosenberg, 2007).
Morellia simplex Loew, 1857
Lipetsk, Donskoe (52.61° N, 38.99° E) env., 23-28.08.1999, K. Tomkovich, 1 S. Samara (Rosenberg, 2007). Voronezh (Negrobov, 2005).
Musca autumnalis De Geer, 1776
Mordovia (Vikhrev et al., 2020). Penza (Lyubvina, 2016b). Samara (Rosenberg, 2007). Voronezh (Negrobov, 2005).
Musca domestica Linnaeues, 1758
Mordovia (Vikhrev et al., 2020). Penza (Levkovich & Levkovich, 1979). Samara (Rosenberg, 2007). Voronezh (Negrobov, 2005).
Musca larvipara Porchinskiy, 1910
Penza (Lyubvina, 2016b). Samara (Rosenberg, 2007). Voronezh (Negrobov, 2005).
Musca osiris (Wiedemann, 1830)
Samara (Lyubvina, 2016a).
Musca tempestiva Fallen, 1817
Mordovia (Vikhrev et al., 2020). Voronezh (Negrobov, 2005).
Muscina angustifrons Loew, 1858
Mordovia (Esin et al., 2023).
Muscina levida Harris, 1780
Lipetsk, Lipetsk, 52.64° N, 39.70° E, Voronezh River, N. Vikhrev, 26.07.2022, 1 S. Mordovia (Vikhrev et al., 2020). Penza (Lyubvina, 2016b).
Nizhny Novgorod, Dzerzhinsk env., 56.21° N, 43.62° E, 18.08.2009, N. Vikhrev, 2 S. Samara (Rosenberg, 2007).
Tambov, W of Kirsanov, 52.65° N, 42.62° E, 27-28.07.2022, N. Vikhrev, 2 S. Tatarstan (Yuzekaeva, 2019). Voronezh (Negrobov, 2005).
Muscina pascuorum Meigen, 1826
Mordovia (Vikhrev et al., 2020).
Samara, Shiryaevo env., 53.40° N 50.05° E, I. Melnik, 13.07.2012, 1 S. Tatarstan (Yuzekaeva, 2019).
Muscina prolapsa (Harris, [1780])
Samara (Lyubvina, 2016a).
Muscina stabulans Fallen, 1817
Chuvashia, Yadrin env., 55.88° N, 46.29° E, N. Vikhrev, 13-15.07.2009, 2 S. Mordovia (Vikhrev et al., 2020).
Penza, Poim env., 53.0° N, 43.2° E, 27.07.2022, N. Vikhrev, 2 S. Samara (Lyubvina, 2016a).
Tambov, W of Kirsanov, 52.65° N, 42.62° E, 27-28.07.2022, N. Vikhrev, 2 S. Tatarstan (Yuzekaeva, 2020).
Voronezh, Khoper SNR, Dubovaya Khata, 51.25° N, 41.78° E, K. Tomkovich,
6.08.2022, 1 S.
Mydaea affinis Meade, 1891
Mordovia (Vikhrev et al., 2020).
Mydaea ancilla Meigen, 1826
Mordovia (Vikhrev et al., 2020).
Mydaea corni Scopoli, 1763
Mordovia (Vikhrev et al., 2020). Samara (Rosenberg, 2007).
Mydaea electa Zetterstedt, 1860
Mordovia (Vikhrev et al., 2020).
Mydaea humeralis Robineau-Desvoidy, 1830
Mordovia (Vikhrev et al., 2020). Samara (Lyubvina, 2016a).
Mydaea nebulosa Stein, 1893
Mordovia (Vikhrev et al., 2020).
Mydaea nubila Stein, 1916
Mordovia (Vikhrev et al., 2020).
Mydaea orthonevra Macquart, 1835
Mordovia (Vikhrev et al., 2020).
Mydaea palpalis Stein, 1916
Penza (Lyubvina, 2016b).
Mydaea setifemur Ringdahl, 1924
Mordovia (Vikhrev et al., 2020). Samara (Rosenberg, 2007).
Mydaea urbana Meigen, 1826
Mordovia (Vikhrev et al., 2020).
Tambov, W of Kirsanov, 52.65° N, 42.62° E, 27-28.07.2022, N. Vikhrev,
3 ?.
Voronezh, Khoper SNR, Dubovaya Khata, 51.25° N, 41.78° E, K. Tomkovich, 6.08.2022, 1
Myospila meditabunda Fabricius, 1781
Mordovia (Vikhrev et al., 2020). Penza (Lyubvina, 2016b). Samara (Lyubvina, 2016a).
Volgograd, Serafimovich env., 49.57° N, 42.60° E, 28-29.04.2013, N. Vikhrev,
2 S.
Voronezh (Negrobov, 2005).
Neomyia cornicina Fabricius, 1781
Mordovia (Vikhrev et al., 2020). Penza (Lyubvina, 2016b). Samara (Rosenberg, 2007). Voronezh (Negrobov, 2005).
Neomyia viridescens Robineau-Desvoidy, 1830
Mordovia (Vikhrev et al., 2020). Voronezh (Negrobov, 2005).
Phaonia aeneiventris Zetterstedt, 1845
Mordovia (Vikhrev et al., 2020).
Phaonia angelicae Scopoli, 1763
Chuvashia, Yadrin env., 55.88° N, 46.29° E, N. Vikhrev, 13.07.2009, 1 Mordovia (Vikhrev et al., 2020).
Ryazan, Tuma env., 55.143° N, 40.616° E, N. Vikhrev, 21.07.2013, 1 Samara (Lyubvina, 2011).
Vladimir, S of Vladimir, 56.05 ° N, 40.41° E, 15.08.2009, N. Vikhrev, 1 Voronezh (Negrobov, 2005).
Phaonia boleticola Rondani, 1866
Tambov, W of Kirsanov, 52.65° N, 42.62° E, 27-28.07.2022, N. Vikhrev, 1 Samara, Enduraykino vill., 53.8° N 51.38° E, 12.08.22, A. Kuzovenko, 3 S,
5 ?.
Ulyanovsk, Shilovka vill. env., 52.82° N, 48.35° E, 1-3.09.2010, K. Tomkovich, 3
Voronezh (Negrobov, 2005).
Phaonia canescens Stein, 1916
Mordovia (Vikhrev et al., 2020).
Phaonia cincta Zetterstedt, 1846
Mordovia (Vikhrev et al., 2020)
Phaonia errans Meigen 1826
Mordovia (Vikhrev et al., 2020). Penza (Lyubvina, 2016b).
Phaonia falleni Michelsen, 1977
Mordovia (Vikhrev et al., 2020).
Lipetsk, Lipetsk, 52.64° N, 39.70° E, Voronezh River, N. Vikhrev, 26.07.2022,
1 ?.
Phaonia fugax Tiensuu, 1946
Voronezh (Negrobov, 2005).
Phaonia fuscata Fallen, 1825
Mordovia (Vikhrev et al., 2020). Samara (Lyubvina, 2016a).
Phaonia gobertii Mik, 1881
Mordovia (Vikhrev et al., 2020).
Phaonia gracilis Stein, 1916
Samara (Rosenberg, 2007).
Phaonia incana Wiedemann, 1817
Mordovia (Vikhrev et al., 2020).
Tambov, W of Kirsanov, 52.65° N, 42.62° E, 27-28.07.2022, N. Vikhrev, 2
Phaonia kowarzii Schnabl, 1887
Mordovia (Vikhrev et al., 2020).
Phaonia laeta Fallen, 1823
Mordovia (Vikhrev et al., 2020).
Phaonia lugubris (Meigen, 1826)
Samara (Lyubvina, 2016a).
Phaonia magnicornis Zetterstedt, 1845
Mordovia (Vikhrev et al., 2020).
Phaonia nymphaearum Robineau-Desvoidy, 1830
Mordovia (Vikhrev et al., 2020).
Phaonia pallida Fabricius, 1787
Mordovia (Vikhrev et al., 2020).
Nizhny Novgorod (Vikhrev & Erofeeva, 2018).
Penza (Lyubvina, 2016b).
Saratov (Vikhrev & Erofeeva, 2018).
Samara (Lyubvina, 2016a).
Tambov, W of Kirsanov, 52.65° N, 42.62° E, 27-28.07.2022, N. Vikhrev, 2 Volgograd (Vikhrev & Erofeeva, 2018). Voronezh (Negrobov, 2005).
Phaonia palpata Stein, 1897
Mordovia (Vikhrev et al., 2020).
Nizhny Novgorod, Vetluzhsky env., 57.2° N, 45.1° E, 11-18.09.2019, A. Matyukhin, 1
Ryazan (Vikhrev & Erofeeva 2017). Samara (Lyubvina, 2016a).
Phaonia rufiventris Scopoli, 1763
Mordovia (Vikhrev et al., 2020).
Vladimir (Vikhrev & Erofeeva, 2018).
Phaonia serva Meigen, 1826
Mordovia (Vikhrev et al., 2020). Penza (Lyubvina, 2016b).
Vladimir, S of Vladimir, 56.05 ° N, 40.41° E, 16.06.2013, N. Vikhrev, 1
Phaonia subventa Harris, 1780
Mordovia (Vikhrev et al., 2020). Ryazan (Vikhrev & Erofeeva, 2018).
Phaonia tiefii Schnabl, 1888
Mordovia (Vikhrev et al., 2020).
Penza, Poim env., 53.0° N, 43.2° E, 27.07.2022, N. Vikhrev, 1 S.
Phaonia tuguriorum Scopoli, 1763
Mordovia (Vikhrev et al., 2020). Voronezh (Negrobov, 2005).
Phaonia valida Harris, 1780
Mordovia (Vikhrev et al., 2020). Samara (Lyubvina, 2016a).
Voronezh, 20 km W of Liski (51.00° N, 39.2° E), K. Tomkovich, 8-9.07.2012,
1 ?.
Phaonia zugmayeriae Schnabl, 1888
Mordovia (Vikhrev et al., 2020).
Polietes domitor Harris, 1780
Mordovia (Vikhrev et al., 2020). Voronezh (Negrobov, 2005).
Polietes lardarius Fabricius, 1781
Mordovia (Vikhrev et al., 2020). Samara (Lyubvina, 2016a). Voronezh (Negrobov, 2005). Polietes steinii Ringdahl, 1913 Mordovia (Vikhrev et al., 2020).
Potamia littoralis Robineau-Desvoidy, 1830
Mordovia (Vikhrev et al., 2020).
Pyrellia rapax Harris, 1780
Mordovia (Vikhrev et al., 2020).
Samara (Rosenberg, 2007). Voronezh (Negrobov, 2005).
Pyrellia secunda Zimin, 1951
Samara (Lyubvina, 2016a).
Pyrellia vivida Robineau-Desvoidy, 1830
Mordovia (Vikhrev et al., 2020). Samara (Lyubvina, 2016a). Voronezh (Negrobov, 2005).
Schoenomyza litorella Fallen, 1823
Mordovia (Vikhrev et al., 2020).
Ryazan, Kasimov env., 54.94° N, 41.34° E, N. Vikhrev, 17-19.07.2013, 1
Spilogona aerea Fallen, 1825
Mordovia (Vikhrev et al., 2020).
Spilogona contractifrons Zetterstedt, 1838
Mordovia (Vikhrev et al., 2020).
Spilogona depressula Zetterstedt, 1845
Mordovia (Vikhrev et al., 2020).
Spilogona pacifica (Meigen, 1826)
Samara (Lyubvina, 2016a).
Spilogona surda Zetterstedt, 1845
Mordovia (Vikhrev et al., 2020).
Stomoxys calcitrans Linnaeus, 1758
Lipetsk, railway station 265th km (52.62° N, 39.47° E), K. Tomkovich, 22.08.1999, 1
Mordovia (Vikhrev et al., 2020). Penza (Lyubvina, 2016b). Samara (Rosenberg, 2007). Tambov (Molev et al., 1967). Voronezh (Negrobov, 2005).
Thricops cunctans Meigen, 1826
Mordovia (Vikhrev et al., 2020).
Thricops diaphanus (Wiedemann, 1817)
Samara (Lyubvina, 2016a).
Thricops lividiventris (Zetterstedt, 1845)
Samara (Lyubvina, 2016a).
Thricops longipes (Zetterstedt, 1845)
Ulyanovsk (Vikhrev, 2010).
Thricops nigrifrons Robineau-Desvoidy, 1830
Mordovia (Vikhrev et al., 2020). Vladimir (Vikhrev, 2010).
Thricops nigritellus Zetterstedt, 1838
Mordovia (Vikhrev et al., 2020). Samara (Rosenberg, 2007).
Thricops semicinereus Wiedemann, 1817
Mordovia (Vikhrev et al., 2020).
Nizhny Novgorod (Vikhrev & Sorokina 2009).
Samara (Rosenberg, 2007).
Thricops simplex Wiedemann, 1817
Mordovia (Vikhrev et al., 2020).
Nizhny Novgorod (Vikhrev & Sorokina, 2009).
Samara (Rosenberg, 2007).
Tambov, W of Kirsanov, 52.65° N, 42.62° E, 27-28.07.2022, N. Vikhrev, 1 S.
Thricops sudeticus Schnabl, 1888
Mordovia (Vikhrev et al., 2020).
FANNIIDAE
Fannia aequilineata Ringdahl, 1945
Voronezh (Negrobov, 2005).
Fannia armata (Meigen, 1826)
Chuvashia (Vikhrev, 2018). Mordovia (Vikhrev et al., 2020). Samara (Lyubvina, 2017).
Fannia barbata Stein, 1892
Mordovia (Esin et al., 2023). Samara (Lyubvina, 2017).
Fannia canicularis Linnaeus, 1761
Chuvashia, Yadrin env., 55.88° N, 46.29° E, N. Vikhrev, 13.07.2009, 2 S.
Lipetsk, Lipetsk, 52.64° N, 39.70° E, Voronezh River, N. Vikhrev, 26.06.2022,
1 S.
Mordovia (Vikhrev et al., 2020). Penza (Levkovich & Levkovich, 1979). Samara (Rosenberg, 2007; Lyubvina, 2017).
Voronezh, Khoper SNR, Varvarino, 51.21° N, 41.72° E, K. Tomkovich, 45.08.2022, 1 S.
Fannia carbonaria (Meigen, 1826)
Samara (Lyubvina, 2017).
Fannia carbonella (Stein, 1895)
Samara (Lyubvina, 2017).
Fannia corvina Verrall, 1892
Mordovia (Vikhrev et al., 2020). Samara (Lyubvina, 2017).
Fannia difficilis (Stein, 1895)
Samara (Lyubvina, 2017).
Fannia fuscula (Fallen, 1825)
Mordovia (Vikhrev et al., 2020).
Tambov, W of Kirsanov, 52.65° N, 42.62° E, 27-28.07.2022, N. Vikhrev, 1 S. Samara (Lyubvina, 2017).
Voronezh, Khoper SNR, Varvarino, 51.21° N, 41.72° E, K. Tomkovich, 45.08.2022, 1 S.
Fannia genualis (Stein, 1895)
Mordovia (Vikhrev et al., 2020).
Fannia hirticeps (Stein, 1892)
Mordovia (Esin et al., 2023). Samara (Lyubvina, 2017). Fannia incisurata Zetterstedt, 1838
Chuvashia, Yadrin env., 55.88° N, 46.29° E, N. Vikhrev, 14-15.07.2009, 3S. Mordovia (Vikhrev et al., 2020).
Fannia latipalpis (Stein, 1892)
Penza (Lyubvina, 2016b).
Fannia lineata (Stein, 1895)
Voronezh (Tewelde, Gaponov, 2019).
Fannia lucidula (Zetterstedt, 1860) (= F. glaucescens)
Nizhny Novgorod, Dzerzhinsk env., 56.21° N, 43.62° E, 16.08.2009, N. Vikhrev, 1
S, 1 ?.
Ryazan, Kasimov env., Oka River, 54.94° N, 41.34° E, N. Vikhrev, 2124.07.2013, 1 S. Oka,1 S. Ranova River, 53.72° N, 39.93° E, N. Vikhrev, 6.06.2012,
1 S.
Fannia lustrator (Harris, 1780)
Mordovia (Vikhrev et al., 2020).
Voronezh, Borisoglebsk (51.4° N, 42.0° E), G. Viktorov, 23.06.1955, 1 Samara (Lyubvina, 2017).
Fannia manicata (Meigen, 1826)
Mordovia (Vikhrev et al., 2020).
Fannia metallipennis Zetterstedt, 1838
Mordovia (Vikhrev et al., 2020).
Fannia minutipalpis (Stein, 1895)
Samara (Lyubvina, 2017).
Fannia monilis Haliday, 1838
Chuvashia, Yadrin env., 55.88° N, 46.29° E, N. Vikhrev, 14-15.07.2009, 5S. Mordovia (Vikhrev et al., 2020).
Fannia ornata (Meigen, 1826)
Samara (Lyubvina, 2017).
Voronezh, Khoper SNR, Dubovaya Khata, 51.25° N, 41.78° E, K. Tomkovich,
8.08.2022, 1 S.
Fannia pallitibia (Rondani, 1866)
Tatarstan (Yuzekaeva, 2019).
Fannia parva (Stein, 1895)
Mordovia (Vikhrev et al., 2020). Samara (Lyubvina, 2017).
Voronezh, Khoper SNR, Varvarino, 51.21° N, 41.72° E, K. Tomkovich, 14.08.2022, 1 S, 1 ?.
Fannia pauli Pont, 1997
Mordovia (Vikhrev et al., 2020).
Fannia polychaeta (Stein, 1895)
Mordovia (Vikhrev et al., 2020).
Tambov, W of Kirsanov, 52.65° N, 42.62° E, 27-28.07.2022, N. Vikhrev, 1 S. Samara (Lyubvina, 2017).
Fannia posticata (Meigen, 1826)
Mordovia (Vikhrev et al., 2020).
Fannia rondanii (Strobl, 1893)
Mordovia (Vikhrev et al., 2020).
Fannia scalaris (Fabricius, 1794)
Chuvashia, Yadrin env., 55.88° N, 46.29° E, N. Vikhrev, 14-15.07.2009, 5S. Lipetsk, Lipetsk, 52.64° N, 39.70° E, Voronezh River, N. Vikhrev, 26.06.2022,
1 S.
Mordovia (Vikhrev et al., 2020). Penza (Levkovich & Levkovich, 1979). Samara (Lyubvina, 2017).
Tambov, W of Kirsanov, 52.65° N, 42.62° E, 27-28.07.2022, N. Vikhrev, 1 S. Tatarstan (Yuzekaeva, 2019).
Voronezh, Khoper SNR, Varvarino, 51.21° N, 41.72° E, K. Tomkovich, 9-13.08.2022, 1 S.
Fannia serena (Fallen, 1825)
Mordovia (Vikhrev et al., 2020).
Chuvashia, Yadrin env. 55.924° N, 46.213° E, N. Vikhrev, 15.07.2009, 1 S. Fannia similis (Stein, 1895)
Chuvashia, Yadrin env., 55.88° N, 46.29° E, N. Vikhrev, 13.07.2009, 2 S. Mordovia (Vikhrev et al., 2020).
Ryazan, Kasimov env., Oka River, 54.94° N, 41.34° E, N. Vikhrev, 21-24.07.2013, 1 S.
Samara (Lyubvina, 2017).
Fannia sociella (Zetterstedt, 1845)
Mordovia (Vikhrev et al., 2020). Samara (Lyubvina, 2017). Fannia spathiophora Malloch, 1918
Mordovia (Vikhrev et al., 2020).
Fannia umbrosa (Stein, 1895)
Mordovia (Vikhrev et al., 2020). Samara (Lyubvina, 2017).
Fannia vespertilionis Ringdahl, 1934
Mordovia (Vikhrev et al., 2020).
Piezura graminicola (Zetterstedt, 1846)
Mordovia (Vikhrev et al., 2020).
Piezura pardalina (Rondani, 1866)
Mordovia (Esin et al., 2023).
SCIOMYZIDAE Anticheta analis Meigen, 1830
Samara, Samara River, Formalny, 53.1° N 50.7° E, 30.07.2020, A. Kuzovenko, 4 6, 1 ?.
Voronezh (Negrobov, 2005).
Anticheta atriseta Loew, 1849
Mordovia (Vikhrev et al., 2020).
Samara, Enduraykino vill., 53.8° N 51.38° E, 12.08.22, A. Kuzovenko, 1 6. Voronezh (Gaponov, 2016).
Anticheta brevipennis (Zetterstadt, 1846)
Voronezh (Gaponov, 2016).
Anticheta nigra (Karl, 1921)
Samara (Rosenberg, 2007).
Colobaea bifasciella Fallen, 1820
Mordovia (Vikhrev et al., 2020).
Ryazan, Kasimov env., Zelenoe, 54.94N 41.34E, N. Vikhrev, 17-19.06.2013, 1 6. Samara, 8 km SE of Kinel, 53.1° N 50.7° E, A. Kuzovenko, 30.07.2020, 1 6, 1 ?. Tambov, Uvarovo env., Vorona river, YPT, M. Esin, 4-6.07.2022, 1 Voronezh (Negrobov, 2005; Gaponov, 2016).
Colobaea distincta Meigen, 1830
Mordovia (Vikhrev et al., 2020).
Samara, 8 km SE of Kinel, 53.1° N 50.7° E, A. Kuzovenko, 30.07.2020, 1 6. Volgograd, Tinguta (48.3° N, 44.3° E), 5.07.1953, G. Viktorov, 1 Voronezh, near Khoper River, 51.36° N, 42.05° E, 7.06.2012, N. Vikhrev, 1 6.
Colobaea pectoralis (Zetterstedt, 1847)
Voronezh (Gaponov, 2016).
Colobaea punctata Lundbeck, 1923
Samara, 8 km SE of Kinel, 53.1° N 50.7° E, A. Kuzovenko, 30.07.2020, 3 6. Voronezh (Negrobov, 2005).
Coremacera catenata Loew, 1847
Samara, Zhiguli (53.4° N, 49.3° E), Novoderzhkin, 3.07.1938, 1 S. Chistovsky, 3.08.1950, 1 $ (ZIN); Rosenberg, 2007. Voronezh (Gaponov, 2016).
Coremacera marginata (Fabricius, 1775)
Lipetsk, railway station 265th km, 52.62° N, 39.47° E, K. Tomkovich, 22.08.1999, 1 S.
Mordovia (Vikhrev et al., 2020).
Ryazan, Kasimov env., Zelenoe, 54.969° N, 41.327° E, N. Vikhrev, 2126.07.2013, 1 $.
Samara (Rosenberg, 2007).
Volgograd, Sarepta, 48.52° N, 44.51° E, T. Bekker, 1868, 1 S, 2 $ (ZIN). Voronezh (Negrobov, 2005).
Dichetophora finlandica Verbeke, 1964
Mordovia (Vikhrev et al., 2020). Penza (Lyubvina, 2016b). Voronezh (Negrobov, 2005).
Dictya umbrarum (Linnaeus, 1758)
Voronezh (Negrobov, 2005).
Ditaeniella grisescens Meigen, 1830
Ulyanovsk (Vikhrev & Murphy, 2022). Voronezh (Gaponov, 2016).
Elgiva cucularia (Linnaeus, 1767)
Mordovia (Vikhrev et al., 2020). Vladimir (Pavlov, 2006).
Voronezh (Negrobov, 2005; Vikhrev & Yanbulat 2021).
Elgiva solicita (Harris, 1780)
Mordovia (Vikhrev et al., 2020).
Penza, Poim env., 53.0° N, 43.2° E, 27.07.2022, N. Vikhrev, 1 S, 1 $. Vladimir (Pavlov, 2006).
Voronezh, Khoper SNR, Varvarino, 51.21° N, 41.72° E, K. Tomkovich, 9-13.08.2022, 3 S.
Euthycera chaerophylli Fabricius, 1798
Mordovia (Dvorak et al., 2020). Nizhny Novgorod (Dvorak et al., 2022). Voronezh (Gaponov, 2016).
Euthycera stictica (Fabricius, 1805)
Voronezh (Gaponov, 2016).
Hydromya dorsalis (Fabricius, 1775)
Vladimir (Pavlov, 2006). Voronezh (Gaponov, 2016).
Ilione albiseta Scopoli, 1763
Ryazan, Khrapovo, 54.61° N, 39.62° E, V. Kovalev, 6.07.1975, 2 S. Samara, Samara, Bostanjoglo, 13.08.1910, 2 $.
Voronezh, Shilovsky forest, 51.58° N, 39.18° E, 7.09.1965, O. Negrobov, 1 $ (ZIN). Ramon, 51.92° N, 39.35° E, A. Shatalkin; 12-15.09.1978, 1 S, 1 $. Voronezh, Khoper SNR, Dubovaya Khata, 51.25° N, 41.78° E, K. Tomkovich,
6.08.2022, 1 S.
Ilione lineata Fallen, 1820
Mordovia (Vikhrev et al., 2020). Voronezh (Gaponov, 2016).
Ilione rossica Mayer, 1953
Ryazan, Ranova River, 53.72° N, 39.93° E, N. Vikhrev, 6.06.2012, 2 S. Samara, Samara, Bostanjoglo, 13.08.1910, 1 $.
Volgograd, Sarepta, 48.52° N, 44.51° E, T. Bekker, 1868, 1 S, 1 $ (ZIN). Voronezh, Ramon, 51.92° N, 39.35° E, A. Shatalkin; 12.09.1978, 6 S, 2 $. Borisoglebsk, 51.37° N, 42.04° E, V. Kovalev, 3.08.1964, 1 $.
Limnia paludicola Elberg, 1965
Voronezh (Gaponov, 2016).
Limnia unguicornis Scopoli, 1763
Chuvashia, Yadrin env., 55.88° N, 46.29° E, N. Vikhrev, 14.07.2009, 1 $. Lipetsk, Lipetsk, 52.64° N, 39.70° E, Voronezh River, N. Vikhrev, 26.07.2022,
2 S.
Mordovia (Vikhrev et al., 2020).
Nizhny Novgorod, Kstov distr., Royka vill., N. Khrustaleva, 18.06.1980, 1 S. Penza (Lyubvina, 2016b).
Tambov, Kirsanov env., Vorona River, 52.61° N, 42.8° E, 28.07.2022, N. Vikhrev, 1 $.
Voronezh, Khoper SNR, Varvarino, 51.21° N, 41.72° E, K. Tomkovich, 14.08.2022, 1 S.
Pelidnoptera nigripennis Fabricius, 1794
Samara (Rosenberg, 2007).
Volgograd: Scherbakovskaya balka (50.49° N, 45.71° E), T. Galinskaya, 25.06.2007, 1 S. Kamyshin (50.1° N, 45.4° E) env., balka, Ig. Melnik, 4.06.2009, 1 S. Kamyshin (50.1° N, 45.4° E) env., balka, G. Viktorov, 17.06.1949, 1 S.
Pherbellia albocostata Fallen, 1820
Mordovia (Vikhrev et al., 2020). Voronezh (Gaponov, 2016).
Pherbellia argyra Verbeke, 1967
Lipetsk, Ryazanka vill., 53.5° N, 39.58° E, A. Yakovlev, 1881, 1 $ (ZIN). Mordovia (Vikhrev et al., 2020).
Voronezh, Ramon, 51.92° N, 39.35° E, A. Shatalkin, 12.09.1978, 1 S, 1 $. Vladimir, S of Vladimir, 56.06° N, 40.46° E, 5.06.2011, N. Vikhrev, 1 S.
Pherbellia austera Meigen, 1830
Mordovia (Vikhrev et al., 2020).
Pherbellia brunnipes Meigen, 1838
Mordovia (Vikhrev et al., 2020). Voronezh (Vikhrev et al., 2020).
Pherbellia cinerella Fallen, 1820
Mordovia (Vikhrev et al., 2020)
Ryazan, Gremyachka, 53.92° N, 39.53° E, Danki uyezd = Skopin distr., A. Semenov, 07.1890, 1 S (ZIN). Ranova River, 53.72° N, 39.93° E, N. Vikhrev, 6.06.2012, 1 $.
Voronezh, Ramon, 51.92° N, 39.35° E, A. Shatalkin, 12.09.1978, 1 S. Chuvashia, Yadrin env., 55.88° N, 46.29° E, N. Vikhrev, 15.07.2009, 2 S.
Pherbellia clathrata Loew, 1874
Samara, Zhiguli, 53.4° N 49.3° E, Novoderzhkin, 4.09.1948, 1 S, 1 $ (ZIN). Volgograd, Elton Lake, mouth of Samaroda River, 49.12N 46.80E, . Astakhov, 17.07.2012, 1 S (ZIN).
Ulyanovsk, Vyazovka env., 52.82° N, 48.35° E, 2-8.05.2011, K. Tomkovich,
1 $.
Pherbellia dorsata (Zetterstadt, 1846)
Voronezh (Gaponov, 2016).
Pherbellia dubia (Fallen, 1820)
Mordovia (Vikhrev et al., 2020). Voronezh (Gaponov, 2016).
Pherbellia griseola (Fallen, 1820)
Lipetsk, Lipetsk, 52.64° N, 39.70° E, Voronezh River, N. Vikhrev, 26.06.2022, 1 S.
Mordovia (Vikhrev et al., 2020).
Ryazan, Kasimov env., Zalesnoe, 54.969° N, 41.327° E, N. Vikhrev, 21-26.07.2013, 4 S, 1 $.
Voronezh, Ramon (51.92° N, 39.35° E), A. Shatalkin, 12.09.1978, 2 S, 4 $.
Pherbellia nana (Fallen, 1820)
Samara (Rosenberg, 2007). Voronezh (Gaponov, 2016).
Pherbellia obtusa (Fallen, 1820)
Mordovia (Vikhrev et al., 2020)
Voronezh, Ramon (51.92° N, 39.35° E), A. Shatalkin, 12.09.1978, 2 S, 4 $. Volgograd, Kalach-na-Donu env., 48.65° N, 43.60° E, 30.04.2013, N. Vikhrev,
1 S.
Pherbellia pallidiventris (Fallen, 1820)
Samara (Rosenberg, 2007). Ulyanovsk (Dvorak et al., 2022). Voronezh (Gaponov, 2016).
Pherbellia pilosa Hendel, 1902
Chuvashia (Borisova et al., 2018). Mordovia (Vikhrev et al., 2020). Ryazan (Vikhrev & Murphy, 2022).
Tambov, Kirsanov env., Vorona River, 52.61° N, 42.8° E, 28.07.2022, N. Vikhrev, 1 S. W of Kirsanov, 52.65° N, 42.62° E, 27-28.07.2022, N. Vikhrev, 1 S.
Voronezh, Khoper SNR, Varvarino, 51.21° N, 41.72° E, K. Tomkovich, 14.08.2022, 1 $. Khoper SNR, Dubovaya Khata, 51.25° N, 41.78° E, K. Tomkovich, 6.08.2022, 2 $.
Pherbellia schoenherri (Fallen, 1826)
Mordovia (Vikhrev et al., 2020).
Ulyanovsk, Radishchevo distr., 52.836N 48.367E, pond, K. Tomkovich, 7-9.05.2011, 1 S.
Vladimir (Pavlov, 2006).
Volgograd, Gulyayevsky, 50.418° N, 42.760° E, N. Vikhrev, 7.07.2012, 2 S. Voronezh, Ramon, 51.92° N, 39.35° E, A. Shatalkin; 12-15.09.1978, 2 S, 1 $.
Pherbellia scutellaris von Roser, 1840
Mordovia (Vikhrev et al., 2020). Voronezh (Gaponov, 2016).
Pherbellia sordida Hendel, 1902
Mordovia (Vikhrev et al., 2020).
Pherbina coryleti Scopoli, 1763
Mordovia (Vikhrev et al., 2020).
Ryazan, Gremyachka, 53.92° N, 39.53° E, Danki uyezd = Skopin distr., A. Semenov: 19.06.1883, 1 S. 8.06.1890, 1 $. 31.05.1908, 1 S (ZIN).
Volgograd, Sarepta, 48.52° N, 44.51° E, T. Bekker, 1868, 1 S, 1 $ (ZIN). Voronezh, Ramon, 51.92° N, 39.35° E, A. Shatalkin; 12.09.1978, 1 S.
Pherbina intermedia Verbeke, 1848
Voronezh (Gaponov, 2016).
Psacadina vittigera Schiner, 1864 (= Psacadina verbekei Rozkosny, 1975)
Mordovia (Vikhrev et al., 2020). Voronezh (Gaponov, 2016).
Psacadina zernyi Mayer, 1953
Mordovia (Vikhrev et al., 2020).
Ulyanovsk Vyazovka env., 52.82° N, 48.35° E, 2-8.05.2011, K. Tomkovich, 1 S. Vladimir (Pavlov, 2006).
Volgograd, Kalach-na-Donu env., 48.65° N, 43.60° E, 30.04.2013, N. Vikhrev,
1 S.
Voronezh, Khoper SNR, Dubovaya Khata, 51.25° N, 41.78° E, K. Tomkovich, 6.08.2022, 1 $.
Pteromicra angustipennis (Staeger, 1845)
Voronezh (Gaponov, 2016).
Pteromicra glabricula (Fallen, 1820)
Mordovia (Vikhrev et al., 2020).
Ryazan, Kasimov env., Zalesnoe, 54.969° N, 41.327° E, N. Vikhrev, 21-26.07.2013, 1 S.
Samara (Rosenberg, 2007). Voronezh (Gaponov, 2016).
Pteromicra leucopeza (Meigen, 1838)
Mordovia (Vikhrev et al., 2020).
Ulyanovsk, Shilovka vill. env., 52.82° N, 48.35° E, 1-3.09.2010, K. Tomkovich, 1 S.
Voronezh (Gaponov, 2016).
Pteromicra oldenbergi (Hendel, 1902)
Voronezh (Gaponov, 2016).
Renocera pallida Fallen, 1820
Mordovia (Vikhrev et al., 2020).
Penza (Lyubvina, 2016b).
Ryazan, Kasimov env, Zalesnoe, 54.969° N, 41.327° E, 21-26.07.2013, N. Vikhrev, 1 S.
Voronezh (Negrobov, 2005).
Sciomyza dryomyzina Zetterstedt, 1846 (= Sciomyza lucida (Hendel, 1902)
Chuvashia, Yadrin env., 55.88° N, 46.29° E, N. Vikhrev, 14.07.2009, 1 $. Voronezh (Negrobov, 2005; Gaponov, 2016).
Sciomyza simplex Fallen, 1820
Voronezh (Negrobov, 2005).
Sciomyza testacea Macquart, 1835
Ryazan, Kasimov env., Zelenoe, 54.969° N, 41.327° E, N. Vikhrev, 21-26.07.2013, 12 S, 9 $.
Voronezh, Khoper SNR, Varvarino, 51.21° N, 41.72° E, K. Tomkovich, 4-5.08.2022, 1 S.
Sepedon sphegea Fabricius, 1775
Mordovia (Vikhrev et al., 2020). Ryazan (Vikhrev & Yanbulat, 2019). Samara (Vikhrev & Yanbulat, 2019). Volgograd (Vikhrev & Yanbulat, 2019).
Voronezh, Khoper SNR, Varvarino, 51.21° N, 41.72° E, K. Tomkovich, 9-13.08.2022, 2 $.
Sepedon spinipes Scopoli, 1763
Mordovia (Vikhrev et al., 2020). Saratov (Vikhrev & Yanbulat, 2019).
Tambov, Kirsanov env., Vorona River, 52.61° N, 42.8° E, 28.07.2022, N. Vikhrev, 2 S, 1 $.
Volgograd (Vikhrev & Yanbulat, 2019).
Voronezh, Khoper SNR, Varvarino, 51.21° N, 41.72° E, K. Tomkovich, 4-5.08.2022, 1 S, 1 $.
Tetanocera arrogans Meigen, 1830
Mordovia (Vikhrev et al., 2020).
Voronezh, Ramon, 51.92° N, 39.35° E, A. Shatalkin; 12.09.1978, 1 $. Tetanocera elata Fabricius, 1781
Mordovia (Vikhrev et al., 2020). Voronezh (Negrobov, 2005).
Tetanocera ferruginea Fallen, 1820
Mordovia (Vikhrev et al., 2020).
Ryazan, Kasimov env., Oka River, 54.94° N, 41.34° E, N. Vikhrev, 21-24.07.2013, 1 S.
Voronezh (Negrobov, 2005).
Tetanocera fuscinervis (Zetterstedt, 1838)
Chuvashia, Yadrin env., 55.924° N, 46.213° E, N. Vikhrev, 15.07.2009, 1 S.
Tetanocera hyalipennis von Roser, 1840
Mordovia (Vikhrev et al., 2020).
Tetanocera montana Day, 1881
Chuvashia, ~ Yadrin, 55.924° N, 46.213° E, willow bush near Sura River, N. Vikhrev, 14.07.2009, 1 S.
Voronezh, Ramon, 51.92° N, 39.35° E, A. Shatalkin; 12-15.09.1978, 3 S.
Tetanocera punctifrons Rondani, 1868
Tambov, W of Tambov, 52.677° N, 41.330° E, N. Vikhrev, 6.06.2012, 1 S.
Tetanocera robusta Loew, 1847
Mordovia (Vikhrev et al., 2020). Voronezh (Negrobov, 2005).
Trypetolimnia rossica Mayer, 1953
Samara (Rosenberg, 2007).
Trypetoptera punctulata Scopoli, 1763
Lipetsk, SNR Galichya Gora, A. Shatalkin, 16.09.1978, 1 $. Mordovia (Vikhrev et al., 2020).
Nizhny Novgorod, W of Nizhny Novgorod, 56.28° N, 43.70° E, birch-pin forest, 12.07.2009, N. Vikhrev, 1 S, 1 $. Penza (Lyubvina, 2016b).
Ryazan, Gremyachka, 53.92° N, 39.53° E, Danki uyezd = Skopin distr., A. Semenov: 13.06.1890, 1 $ (ZIN). Kasimov env., Zalesnoe, 54.969° N, 41.327° E, N. Vikhrev, 21-26.07.2013, 1 $.
Samara, Zhiguli, 53.4° N, 49.3° E, Chistovsky, 3-10.08.1948, 1 S, 1 $ (ZIN). Volgograd, Sarepta, 48.52° N, 44.51° E, T. Bekker, 1868, 1 S, 1 $ (ZIN) Voronezh, Kalach, 50.4° N, 41.0° E, I. Gudim, 4.07.1933, 1 S (ZIN). Voronezh SNR, 51.9° N, 39.4° E, Keleynikova, 12-24.06.1957, 1 S, 1 $. Borisoglebsk, 51.4° N, 42.0° E, G. Viktorov, 21.06.1955, 1 S.
Discussion
The table presents information on biodiversity of Diptera from 9 families (Tipulidae, Limoniidae, Pediciidae, Mycetophilidae, Keroplatidae, Ptychopteridae, Muscidae, Fanniidae, Sciomyzidae) inhabiting the territory of 14 regions. The most
studied and diverse fauna is in the Republic of Mordovia. It is represented by 399 species. The fauna of these families is least known from the Republic of Tatarstan, Lipetsk Region, Nizhny Novgorod Region and Vladimir Region (respectively, 9, 19, 20 and 21 species).
Table. The number of species of di ferent families of diptera in individual regions
Regions Tipulidae Limoniidae Pediciidae Mycetophilidae Keroplatidae Ptychopteridae Muscidae Fanniidae Sciomyzidae Total
Chuvashia 37 27 0 5 0 0 14 7 5 95
Lipetsk 0 4 0 0 0 1 7 2 5 19
Mordovia 43 77 6 51 9 2 144 26 35 399
Nizhny Novgorod 0 2 0 0 0 0 14 1 3 20
Penza 0 1 0 0 0 0 35 3 5 44
Ryazan 0 17 1 0 0 0 14 2 14 48
Samara 8 2 0 0 0 0 76 18 18 122
Saratov 21 19 0 0 0 3 6 0 1 50
Tambov 0 11 0 0 0 0 16 3 5 35
Tatarstan 0 0 0 0 0 2 5 2 0 9
Ulyanovsk 25 8 1 5 0 0 13 0 6 58
Vladimir 0 2 0 0 0 0 13 0 6 21
Volgograd 0 8 0 1 1 1 18 0 12 40
Voronezh 4 2 0 5 0 1 50 8 55 125
Total 64 102 6 69 10 4 190 36 64 545
A total of 545 species are known from 9 families of Diptera in the large macroregion (Volga Upland and Oka-Don Plain). The largest number of species include Muscidae (190) and Limoniidae (102). The least known biodiversity includes Ptychopteridae (4 species), Pediciidae (6 species) and Keroplatidae (10 species).
Conclusion
The fauna of these Diptera families is poorly studied in the vast majority out of the 14 regions located within the Volga Upland and Oka-Don Lowland. It is assumed that the fauna of considered families is far from being fully studied. This publication serves as a starting point for further research into the biodiversity of the macroregion.
Acknowledgement
M.O. Yanbulat provided important assistance in compiling the list of Sciomyzidae. The authors are grateful for providing a map of the territories to A.A. Khapugin (Tyumen). This research was funded by the Russian Science Foundation, grant number 22-14-00026. We thank Olga Ovchinnikova and Galina Suleymanova (Saint Petersburg) for the opportunity to examine the important material in ZIN.
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СПИСОК НЕКОТОРЫХ МАЛОИССЛЕДОВАННЫХ СЕМЕЙСТВ DIPTERA ИЗ РЕГИОНОВ ПРИВОЛЖСКОЙ ВОЗВЫШЕННОСТИ И ОКСКО-ДОНСКОЙ НИЗМЕННОСТИ
А.Б. Ручин1*, Н.Е. Вихрев2, Д.И. Гаврюшин2, М.Н. Есин1
1 Объединенная дирекция Мордовского государственного природного заповедника имени П. Г. Смидовича и национального парка «Смольный», Россия
2Зоологический музей МГУ им. М.В. Ломоносова, Россия
1 2 e-mail: '*ruchin.alexander@gmail.com, nikita6510@ya.ru
Diptera - это один из наиболее разнообразных по числу видов отряд среди насекомых. В статье представлены сведения о биоразнообразии Diptera из 9 семейств (Tipulidae, Limoniidae, Pediciidae, Mycetophilidae, Keroplatidae, Ptychopteridae, Muscidae, Fanniidae, Sciomyzidae), обитающих на территории 14 регионов, находящихся в пределах Приволжской возвышенности и Окско-Донской низменности. Наиболее изученная и разнообразная фауна в Республике Мордовия (399 видов). Наименее известна фауна этих семейств из Республики Татарстан, Липецкой области, Нижегородской области и Владимирской области (соответственно, 9, 19, 20 и 21 вид). В общей сложности на территории большого макрорегиона из 9 семейств Diptera известно 545 вида. Наибольшее число видов включают семейства Muscidae (191) и Limoniidae (102). Наименее известно биоразнообразие Ptychopteridae (4 вида), Pediciidae (6 видов) и Keroplatidae (10 видов).
Ключевые слова: Tipulidae, Limoniidae, Pediciidae, Mycetophilidae, Keroplatidae, Ptychopteridae, Muscidae, Fanniidae, Sciomyzidae, Европейская Россия
