The lithostratigrahy and sedimentology of the fossiliferous Upper Cretaceous strata exposed in the Gobi Desert of Mongolia at Ukhaa Tolgod are described and mapped on aerial photos. Topographic features are also mapped by plane table and alidade.
Five lithologic and sedimentologic facies are described: E-1, distinctly cross-stratified sandstone with fine structure, interpreted to represent eolian dune deposits; E-2, vaguely bedded sandstone with cross-stratified concretionary sheets, interpreted to represent eolian dune deposits modified by diagenetic formation of slope-parallel concretionary sheets of pedogenic calcite; S, structureless sandstone lacking concretions or cross-strata, interpreted to represent sandslide deposits generated by mass wasting along the lee slopes; C, conglomerate interpreted to represent basin-margin conglomerates washed into the dune field from adjacent topographic highs; and M, mudstone and siltstone interpreted to represent interdune deposition in ephemeral ponds and lakes. Facies E-2 and S have not been reported previously. Eleven stratigraphic sections at various localities within the Ukhaa Tolgod drainage basin are documented. The exposed composite section consists of about 75 m of pale orange sandstones, greenish-brown conglomerates, and brown siltstones that are products of an arid environment. Four schematic cross sections are documented to illustrate the lateral relationships among the five facies.
In the Ukhaa Tolgod area, the beds dip about 2.5° to the south, away from the nearby Gilbent Range. This structural attitude is interpreted to be related to the uplift of the Gilbent block along normal faults exposed at the base of the range.
The dune-derived sandslides of Facies S contain a rich skeletal fauna of Late Cretaceous dinosaurs, mammals, and lizards. Essentially, all the skeletal remains collected at Ukhaa Tolgod come from Facies S. Facies E-1 does contain numerous, concave-up depressions in the cross-strata interpreted as vertebrate tracks. Facies E-2 contains abundant cylindrical structures interpreted as burrows.
The strata at Ukhaa Tolgod are referred to the Djadokhta Formation. As seen in the Bayn Dzak Member at Bayn Dzak, facies E-1, E-2, S, and M dominate the lower part of the section at Ukhaa Tolgod, with prominent beds of Facies C exposed near the top. Accordingly, the exposures at Ukhaa Tolgod are referred to the Bayn Dzak Member of the Djadokhta Formation. Classic exposures of the Barun Goyot Formation at Khulsan differ in having units of flat-bedded sandstone intercalated with beds of Facies S near the top of the section.
To date, over 1,000 vertebrate skulls and skeletons have been collected from Facies S. Most are preserved as float contained in small calcareous nodules; however, some were found in situ. Many specimens represent either fairly complete skulls or skulls with articulated or associated postcranial skeletons. Based on faunal similarities between Bayn Dzak and Ukhaa Tolgod, the fauna at Ukhaa Tolgod is interpreted to reflect a Campanian age. The rich assemblage of fossils makes Ukhaa Tolgod one of the richest Late Cretaceous vertebrate fossil localities in the world, and the fossils provide unique insights into evolutionary developments of mammals, lizards, and dinosaurs, including birds, less than 10 my before the terminal Cretaceous extinction event.
Introduction
Dashzeveg et al. (2005) provide a summary of research carried out in the classic exposures of the Djadokhta Formation at Bayn Dzak ( = Bayan Zag of Benton, 2000) in the southern Ulan Nur Basin of Mongolia's Gobi Desert. The same authors divide the Djadokhta Formation into a lower Bayn Dzak Member exposed at Bayn Dzak and an upper Tugrugyin Member exposed at Tugrugyin Shireh ( = Tögrögiin Shiree of Benton, 2000). Here, we focus on previous work done in the Nemegt Basin, which lies to the south of the Ulan Nur Basin near Mongolia's border with the People's Republic of China.
Expeditions by the Soviet Academy of Sciences between 1946 and 1949 extended paleontologic exploration in Mongolia southward into the Upper Cretaceous Nemegt Formation of the Nemegt Basin, where they discovered several Latest Cretaceous taxa including Tarbosaurus baatar and Saurolophus angustrirostris (Efremov, 1949, 1954; Maleev, 1952, 1954).
Between 1963 and 1971, the Polish-Mongolian Palaeontological Expeditions discovered rich, Late Cretaceous fossil localities in the Barun Goyot Formation ( = Baruungoyot in Benton, 2000) of the Nemegt Basin to the southwest of Bayn Dzak, as well as the red beds of Khermeen Tsav ( = Hermiin Tsav of Benton, 2000) in the western Nemegt Basin (Kielan-Jaworowska and Dovchin, 1969; Kielan-Jaworowska and Barsbold, 1972). The expeditions assembled impressive collections of Cretaceous mammals (Kielan-Jaworowska, 1969, 1970, 1974, 1975, 1984; see also Kielan-Jaworowska et al., 2000; Kielan-Jaworowska et al., 2003 and 2004, for summaries). The rich fauna of dinosaurs from the Djadokhta and Barun Goyot Formations has been described by Maryańska and Osmólska (e.g., Maryańska, 1971; Maryańska and Osmólska, 1975; Osmólska, 1972, 1976, 1980, 1993). Lizards from these formations are discussed in (e.g., Sulimski, 1975; Borsuk-Bialynicka 1984, 1991a, 1991b; see also Alifanov, 2000, as well as Gao and Norell, 2000, for other faunal summaries).
Soviet-Mongolian (subsequently Russian-Mongolian) expeditions were conducted in Mongolia beginning in 1970, and continued until recently, as reviewed by Kurochkin and Barsbold (2000).
From 1992 to the present, the Japan-Mongolia Joint Paleontological Expedition (JMJPE) has conducted extensive field seasons throughout the Gobi Desert. Recent accounts of their activities are presented in Ishii et al. (2000, 2004).
In 1991, following a brief reconnaissance trip in 1990, the Mongolian Academy of Sciences–American Museum of Natural History Expedition (MAE) began field investigations based on the Agreement on Scientific Cooperation between those two institutions. In 1993, 70 years after Andrews' Central Asiatic Expedition discovered the first well-documented clutches of dinosaur eggs, the first-known skeletons of the dinosaurs Protoceratops and Velociraptor, and the first well-preserved skulls of Late Cretaceous mammals at Bayn Dzak (Osborn, 1924; Simpson, 1925; Gregory and Simpson, 1926; Berkey and Morris, 1927; Andrews, 1932; Morris, 1936), members of the MAE discovered a new fossil locality in the Nemegt Basin called Ukhaa Tolgod, which means “Brown Hills” (Dashzeveg et al., 1995). This locality has yielded an unmatched abundance of well-preserved vertebrate fossils, including the highest concentration of mammalian skulls and skeletons from any known Mesozoic site. More than 100 skeletons of theropod, ankylosaurian, and protoceratopsian dinosaurs have been identified, including specimens of Late Cretaceous birds (Chiappe et al., 1998, 2001, 2002; Schweitzer et al., 1999; Norell and Clarke, 2001; Suzuki et al., 2001), oviraptorid dinosaurs (Clark et al., 2001), including a specimen positioned on top of a nest of eggs in brooding posture (Norell et al., 1995; Clark et al., 1999), a partial ornithomimid braincase (Makovicky and Norell, 1998) and the first known embryo of a theropod (Norell et al., 1994). Johnston et al. (1996) discussed alleged vertebrate eggs that they identified as fossilized pupal chambers. Recent publications regarding the mammalian fauna include Horovitz (2000); Kielan-Jaworowska (1998); Kielan-Jaworowska et al. (2000); Novacek et al. (1997); Rougier et al. (1996, 1997, 1998); Wible and Rougier (2000), and Wible et al. (2004). In all, over 1,000 skulls, many with associated skeletons, of mammals, lizards, and dinosaurs have been identified. In relation to other Mesozoic localities, the concentration and diversity of mammals, lizards, and dinosaurs are unusually high.
As a result of these paleontological expeditions, an extensive body of literature has been published regarding the geology of the Djadokhta and Baurn Goyot formations (Gradziński, 1970; Lefeld, 1965, 1971; Gradziński and Jerzykiewicz, 1972, 1974a, 1974b; Jerzykiewicz, 1998, 2000; Jerzykiewicz and Russell, 1991; Jerzykiewicz et al. 1993a, 1993b; and Shuvalov, 2000).
Large-scale cross-bedded strata of the Djadokhta have been previously interpreted as both deltaic (Tverdochlebov and Tsybin, 1974) and eolian (Gradziński and Jerzykiewicz, 1974b; Eberth, 1993; Jerzykiewicz et al., 1993a, 1993b; Fastovsky et al., 1997). Most recent interpretations have favored an eolian origin for these large-scale cross-stratified sandstones (Eberth, 1993; Fastovsky et al., 1997; Loope et al., 1998). However, the skeletally fossiliferous units at Ukhaa Tolgod are not cross-stratified (Loope et al., 1998). We recognize three different sandstone facies at Ukhaa Tolgod. Two are eolian, containing distinctly to vaguely cross-stratified sandstone. These accumulated during relatively rapid and slow migration of large bedforms. The third and only skeletally fossiliferous sandstone facies is structureless and often contains lenses of granules and pebbles as well as floating cobbles. It has previously been suggested that these rocks also resulted from very rapid eolian deposition (Eberth, 1993). However, Loope et al. (1998, 1999) interpret this facies to represent mass flows that buried lee slopes and (with intermixed fluvial deposits) filled interdunes during mesic intervals when the dunefield was stabilized.
Within the distinctly cross-stratified eolian sandstone are numerous deformed zones that represent vertical cross sections of dinosaur tracks—the first such tracks known from the Late Cretaceous rocks of the Gobi Desert and clear evidence that dinosaurs traversed actively migrating dunes (Loope et al., 1998). Bioturbation and accumulation of clay and pedogenic carbonates in the fossiliferous eolian facies support the sandslide interpretation of the laterally adjacent structureless sandstones. Together, these lines of evidence indicate that the large dunes of the Late Cretaceous landscape were stabilized for long periods of time, and, when active, migrated by deposition of wind-ripple strata and thin grain flows (Hunter, 1977).
Here, in addition to documenting and describing the strata and facies exposed at Ukhaa Tolgod, we evaluate the question of whether these strata should be assigned to the Djadokhta or Barun Goyot Formation and discuss the age of the vertebrate fossils from the site.
Modern Topographic Features and Major Fossil Localities
Ukhaa Tolgod lies about 650 km southwest of Ulaanbaatar in the Nemegt Basin of Mongolia, a post-Oligocene tectonic depression controlled by normal faulting (Gradziński et al., 1977). The Gilbent Uul forms the northern edge of the Nemegt Basin in the area near Ukhaa Tolgod.
Ukhaa Tolgod means “brown hills” in Mongolian, and the field area is marked by a series of reddish brown bluffs, buttes, and escarpments. Overall, the collecting area is about 2.2 km across and follows the bluffs of a northeast-southwest trending drainage basin eroded into the southward-inclined alluvial apron of the Gilbent Uul. It is located at about 43°32′ N latitude and 101°34′ E longitude, and about 1500 m above sea level.
One of us (C. M.) constructed a physiographic map of the Ukhaa Tolgod area using a plane table and alidade (map 1), which documents the locations of eight separate collecting areas in the northern half of the Ukhaa Tolgod drainage basin: Ankylosaur Flats, Sphinx, Sugar Mountain, Death Row, Camel Humps, Xanadu, First Strike, and Zofia's Hill. The facies maps (maps 2–Map 3Map 4Map 56) further document the location of these eight localities, as well as four additional ones in the southern half of the Ukhaa Tolgod drainage basin: Romeo and Juliet, Delta Force, Granger Hill, and Downlap Hill.
Lithologic Units and Sedimentologic Facies
The strata exposed at Ukhaa Tolgod consist of five distinct lithologic and sedimentologic facies, all nonmarine sandstones, siltstones, and conglomerates. Three sandstone facies dominate. Conglomerate is relatively rare, while mudstones and siltstones are thin and laterally restricted.
Eolian Facies
Distinctly Cross-stratified Sandstone with Fine Structure (Facies E-1)
Description
This facies is composed predominantly of moderate reddish orange (10R 6/6) to pale red (10R 6/2) to moderate orange pink (10R 7/4), fine- to coarse-grained sand that was deposited in distinct laminae that consistently dip 25 degrees to the northeast (fig. 1). Nearly all of the distinct cross-strata present at Ukhaa Tolgod lie within one laterally discontinuous set that is typically about 12 m thick. Small-diameter burrows (4–10 mm) and occasional rhizoliths (≤ 2 mm in diameter) are present within these cross-strata. Concave up folds that are circular to oval in plan and range from 10–50 cm across are common along certain cross-strata (fig. 2). Sharp contacts separate the folded from the overlying, unfolded strata, and coarse-grained sediment typically fills the central shaft at the core of each fold. Skeletal fossil remains are not present. On the facies maps (maps 2–6), this facies is represented by Ke.
Interpretation
Large-scale cross-strata in the Late Cretaceous redbeds of the Gobi have been interpreted as eolian by Gradzinski and Jerzykiewicz (1974b) at Bayn Dzak, Jerzykiewicz et al. (1993a, 1993b) and Eberth (1993) at Bayan Mandahu in Inner Mongolia, and Fastovsky et al. (1997) at Tugrugyin Shireh, and we concur with their interpretation (Loope et al., 1998, 1999). Some earlier interpretations suggested a deltaic origin for these cross-strata (Tverdochlebov and Tsybin, 1974); however, that interpretation does not conform well with the following evidence.
By making direct observations during sandstorms within the dunefields of coastal Oregon and Texas, Hunter (1977) demonstrated that distinct, relatively thin laminae and beds are produced during the migration of eolian dunes and ripples. The laminae produced by climbing wind ripples are inverse graded. Deposits of extensive Mesozoic and Paleozoic sand seas of the western United States (Kocurek, 1988) are clearly composed of the characteristic bedding types described by Hunter (1977). Such inverse-graded ripple laminations are present but usually poorly preserved at Ukhaa Tolgod. They are clearly present and well developed in laterally equivalent exposures that crop out at Little Ukhaa—a locality about 10 km northwest of Ukhaa Tolgod proper (fig. 3). This distinctly cross-stratified facies (E-1) thus has sedimentary structures similar to those in Holocene dunes and well-studied ancient eolian sandstones from the western U.S. and can thus be confidently interpreted as eolian. Although it has not yet proven possible to discern the velocity of paleowinds from eolian strata, if violent windstorms have left a sedimentary record in Mongolia and the western U.S., this distinctly stratified facies is that record (rather than the structureless sandstone facies discussed below).
Despite the presence of abundant fossil skeletons, dinosaur tracks have only recently been reported from the Djadokhta Formation (Loope et al., 1998). The scarcity of bedding plane exposures and the poor induration of the sandstones mitigate against recognition and preservation of tracks (Jerzykiewicz et al., 1993a, 1993b). Nevertheless, the thin-bedded, easily deformed sediments of this facies accumulated in an environment that was readily accessible to large animals. We confidently interpret the concave-up downfolds seen in vertical cross section in this facies as the tracks of large vertebrates (Loope et al., 1998). Like many tracks in the Jurassic Navajo Sandstone of southern Utah (Loope, 2006), these tracks are represented by smooth folds, and were probably emplaced in dry, cohesionless sand.
Vaguely Bedded Sandstone with Cross-stratified Concretionary Sheets (Facies E-2)
Description
Moderate reddish orange (10R 6/6) to moderate orange pink (10R 7/4), fine to coarse sand nearly devoid of visible depositional structures is cemented by tabular calcareous concretions that are 5–10 cm thick and dip 25 degrees to the northeast—parallel to the cross-strata of Facies E-1. In weathered horizontal exposures, the concretionary sheets weather into strong relief, defining straight to gently arcing ridges (fig. 4). The concretions delineate “sets” analogous to those of cross-strata that are typically in excess of 10 m thick; the thickest set we measured near Delta Force, the base of which was not exposed, is 24.5 m thick.
Reddish yellow (YR 6/6), uncemented sand fills abundant simple, nonmeniscate burrows that cut lighter (7.5 YR7/4) concretions. In thin section, burrow margins are sharp, and calcite ranges from micrite to spar. Sand grains bear thin, often incomplete clay coats that are thickest in depressions in grain surfaces. Rhizoliths and other fossils are absent. On maps 2–Map 3Map 4Map 56, this facies is represented by Ke.
Interpretation
Similarities in the texture and scale between the vague cross-strata of this facies and E-1 allow confident interpretation of Facies E-2 as eolian dune deposits. Although E-2 is nearly devoid of visible depositional structures, the postdepositional development of the concretionary sheets indicates preservation of a slope-parallel sedimentary fabric. At least two interpretations are possible: (1) First, distinct depositional structures were never present because plants acted as sediment baffles. According to Kocurek and Nielson (1986), plants interfere with the propagation of wind ripples, thereby preventing the development of distinctly laminated deposits typically produced by wind ripple lamination and climb. Unless plant cover is extremely uniform, however, baffling of saltating sand by plants should lead to growth of hummocks or plant mounds, the geometry of which is not reflected in the tabular nature of the concretionary sheets in E-2. (2) Alternatively, depositional structures were produced but were destroyed before burial. Although invertebrate burrows that were emplaced relatively deep in the sediment are present in great abundance, the retention of the crude fabric element that repeats at 10–15 cm intervals suggests that the sediments were not deeply “stirred”. Trampling by dinosaurs could have sufficiently mixed the surficial sediment to a depth of 10–15 cm, thereby preventing the preservation of physical structures but allowing the grain-size differences between different depositional packages to be preserved. Although our “dinoturbation” (Lockley and Gillette, 1989) hypothesis for E-2 is speculative, it is consistent with the facts that (1) dinosaur tracks are common in E-1 and (2) that no facies similar to E-2 has been described from well-studied Mesozoic and Paleozoic eolianites that lack vertebrate skeletal remains.
The orientation of the clay that coats the sand grains (parallel to grain surfaces), the presence of clay at grain contacts, the tendency of clay coats to thicken over embayments in the sand grains, and the absence of meniscus bridges between grains indicate that most of the clays within and adjacent to the concretions are inherited clay rims as described by Wilson (1992). Thus the clays were deposited on the grains prior to their arrival at the site of deposition. Such clays record the reworking of grains that were coated by infiltration of clay-charged water and are common in Quaternary dune sands (Wilson, 1992; Loope and Dingus, 1999). Although the postalluvial burrowing could have destroyed meniscus bridges produced by in situ clay accumulation, the absence of these bridges in the burrow fills of the concretions suggests that this process was not volumetrically important at the depth of concretion development.
Eberth (1993) demonstrated that pedogenic calcite is widespread in structureless sandstones of the Djadokhta Formation at Bayan Mandahu in Inner Mongolia but interpreted the calcite cement of cross-stratal toesets as groundwater calcrete. In contrast, the burrows that cut the sheetlike calcareous concretions of E-2 clearly postdate the onset of carbonate accumulation. The cross-cutting burrows, the location of concretions within the sediments deposited high on the slopes of large bedforms (far above the water table), and the fine grain size of the calcite indicate that concretion development started within the vadose zone. Pedogenic calcite plays a pivotal role in our hypothesis explaining the preservation of abundant, articulated, and unscavenged vertebrate skeletons at Ukhaa Tolgod (see Sandslide Facies, below).
A recent study (Yang et al., 2003) of the Badain Jaran Desert of western Inner Mongolia, China, provides a Quaternary analog for the caliche zones that we have recognized in the Cretaceous strata at Ukhaa Tolgod. Yang et al. (2003) interpret the many calcite cemented zones that lie parallel to cross-strata within these dunes as evidence that dune migration ceased during episodes of climate amelioration. The cemented zones are generally 8–10 cm thick. Soil-forming processes were terminated when arid conditions returned and dunes became mobile. In some dunes as many as two dozen cemented zones are present.
The abundance of biogenic structures and grain-rimming clays, plus the presence of pedogenic calcite indicate that the large eolian bedforms upon which these sands accumulated were immobile for at least hundreds, and probably thousands of years. In comparison, the Late Quaternary Nebraska Sand Hills where dunes have been stabilized by vegetation for at least several hundred years, sands are coated by infiltrated clays, but no calcium carbonate has accumulated (Lewis, 1990).
Given the presence of early diagenetic, vadose calcite in this facies, the scarcity of rhizoliths is surprising. Could Late Cretaceous plants survive on dunes? Calcitic rhizoliths are abundant in numerous ancient eolianites (Esteban and Klappa, 1983; Loope, 1988), including the Djadokhta Formation at Bayn Dzak (Loope and Dingus, 1999). Magdefrau (1956) describes a possible example of Early Cretaceous plants that grew on dunes, but the abundant rhizoliths preserved in Paleozoic and Mesozoic eolianites of the Colorado Plateau (all of which are pre-Cretaceous) are developed along horizontal unconformities and do not represent the roots of dune-dwelling plants (Loope, 1988). The large rhizoliths in eolian strata at Bayn Dzak (Loope and Dingus, 1999) appear to record the presence of dune plants—their roots lie parallel to cross-strata (an advantage only within the vadose zone) and laterally adjacent fluvial deposits suggest that streams flowed between contemporaneous dunes.
Sandslide Facies
Structureless Sandstones Lacking Oriented Concretions or Cross-strata (Facies S)
Description
The third sandstone facies, unlike the two described above, lacks physical sedimentary structures and any indicators of a preserved slope-parallel fabric (fig. 5A, B). Isolated or lenticular deposits of granules, pebbles, and cobbles are occasionally present. This facies is the most fossiliferous at Ukhaa Tolgod. Oviraptorid skeletons preserved on nests of well-arranged eggs, along with perfectly articulated skeletons of tiny mammals and lizards are present. At one locality (Delta Force), a 20 m thick body of well-sorted sandstones of facies S abuts the depositional pinchout of eolian cross-strata of facies E-2 that dip at 25 degrees and are at least 24 m thick. At other localities, such as Downlap Hill, the sloping upper surfaces of well-sorted sandstone bodies of facies S are downlapped by eolian cross-strata of facies E-1.
On maps 2–Map 3Map 4Map 56, the structureless sandstone (Ks) is occasionally subdivided into exclusively dune-derived sandstone (Ksd) and crudely flat-bedded sandstone (Ksf), which often contains lenses of larger clasts representing sediments that have washed in from outside the dune field.
Interpretation
Based primarily on taphonomic evidence that supports rapid accumulation and the presence of large scale cross-strata within the formation, numerous authors have interpreted structureless Djadokhta sandstones as products of violent sandstorms (e.g., Eberth, 1993). Several lines of evidence lead us to interpret these deposits as dune-sourced sandslides (Loope et al., 1998, 1999).
Perfect preservation of articulated small lizard and mammal skeletons in this facies strongly suggests that its structureless nature is depositional and not the result of bioturbation. Our sandslide hypothesis is based primarily on the lack of depositional structures and the stratal relationships with facies E-1 and E-2. Somewhat similar deposits have been described from stabilized Quaternary dunefields in the Sahel of Africa and the Great Plains of North America (Talbot and Williams, 1978, 1979; Ahlbrandt and Fryberger, 1981; Sweeney and Loope, 2001). Climate change, dune stabilization, and the attendant accumulation of calcite in the dune deposits led to mass movements that built fans (Loope et al., 1999). Sandslide deposits were downlapped and buried when dunes reactivated.
We cannot cite a Holocene example of animals overwhelmed by mass flows from dunes, and until the publication by Yang et al. (2003; see above) we were also unaware of a dunefield in which dune lee slopes are underlain at shallow depth by calcite-cemented sand. Positive pore water pressure within steeply sloping unconsolidated material is a well-known trigger for mass wasting events (Graham, 1984). Dunefields that lack slope-parallel barriers to the flow of infiltrating water are not likely to generate large mass flows. Thus, many unusual conditions were required to generate the fossiliferous sediments of the Djadokhta Formation: (1) large dunes formed in a subsiding sedimentary basin; (2) dunes became stabilized; (3) dust flux, average rainfall, and duration of dune stability allowed development of calcite zones beneath dune lee slopes; (4) sufficient food and water were available to attract large animals to the site; (5) rare, heavy rainfall events caused perched water tables to form under dune slopes. Yang (personal commun., August, 2003) notes that rain-triggered sandslides are common on the lee slopes of large dunes in western China.
The crudely flat-bedded sandstone (Ksf) was likely deposited by ephemeral streams entering the dunefield from adjacent highlands. Possible modern analogs are the mass flows and hyperconcentrated flows generated by the breaching of dune dams by floods entering the Namib sand sea in southwestern Africa (Svendsen et al., 2003).
Conglomerates (Facies C)
Description
Conglomerates composed primarily of igneous clasts ranging in size from granules to boulders up to 85 cm in diameter are occasionally interbedded with the sandstone facies described above (fig. 6). The igneous clasts are grayish green (10 GY 5/2) to dark yellowish green (10 GY 4/4) and consist of an andesitic matrix with phenocrysts of potassium feldspar. Clasts range in shape from angular to moderately well rounded. Usually these conglomeratic beds are distinct from other units; however, lenses of similar conglomeratic material are occasionally contained in the sandslide deposits of facies S. Vertebrate fossils are absent. On the facies maps 2–Map 3Map 4Map 56, this facies is represented by Kc.
Interpretation
The conglomerates clearly indicate the presence of a topographic high near the Ukhaa Tolgod area. The angular nature of numerous clasts indicates that transport distance from the topographic high to the Ukhaa Tolgod dunefield was not great. The orientation of channels containing the conglomerates suggests that current direction was from northeast to southwest. The conglomerates were probably deposited on alluvial fans that built out from the topographic high and impinged on the dunefield, which probably occupied the central area of the paleobasin. Similar topographic regimes have been reported from other areas in the Gobi, including Bayan Mandahu in northern China (Eberth, 1993).
Interdune Facies
Mudstones and Siltstones (Facies M).
Description
Moderate reddish-brown (10 R 4/6) to dark reddish-brown (10 R 3/4) siltstone contains small amounts of clay and often contains a substantial fraction of fine sand (fig. 7). The siltstone weathers into small blocks that break apart easily. Rip-up clasts are occasionally present. The siltstone beds can be as much as 20 cm thick; however, the beds are not laterally continuous over large geographic areas. Based on mapping and correlations, the most extensive set of interdune layers extends across a one square km area in the eastern portion of the Ukhaa Tolgod Basin. Contacts with over- and underlying units are sharp. Remains of vertebrates are rare. On maps 2–Map 3Map 4Map 56, this facies is represented by Km.
Interpretation
The siltstones appear to represent sediments deposited at the bottom of small ephemeral ponds or lakes that formed when corridors between the dunes at Ukhaa Tolgod were flooded. Although skeletal remains of vertebrates are rare, two fossil turtle shells with partial postcrania have been collected from these deposits, suggesting that the ponds did occasionally serve as sources of water for some of the vertebrates at Ukhaa Tolgod.
Map of Sedimentary Facies
The geologic units exposed in the Ukhaa Tolgod drainage basin were mapped in 1997 on air photos taken at the end of the 1996 field season (maps 2–Map 3Map 4Map 56). The units mapped are labeled in the legend on the index map (map 2). Major fossil localities are plotted by name on the maps. The locations of individual fossil specimens are plotted according to taxon, as specified by symbols listed in the legend. The distributions of the five sedimentary facies found at most localities between Downlap Hill in the southwest and Zofia's North in the northeast are discussed below in Stratigraphic Sections and Schematic Cross Sections.
Across the Ukhaa Tolgod drainage basin on the southern ridge, a series of poorly exposed patches of Facies S and E-1 is capped by Quaternary pediment. One dinosaur nest has been collected from the southernmost patch of Facies S. This locality is named Uranus.
At Xanadu SW, a fault trending essentially east-west cuts the exposures uplifting the southern block and down-dropping the northern one. The southern ridge between Xandu SW and First Strike is capped by Quaternary pediment.
Midway between Camel Humps and First Strike, three small outcrops of cross-stratified dune sand cap the southwest end of the northern edge of the basin. The middle one is cut by a small, north-south trending fault, with the eastern block uplifted and the western block down-dropped.
Stratigraphic Sections and Schematic Cross Sections
The vertical and lateral stratigraphic distributions of the five facies were documented in 11 stratigraphic sections and four schematic cross sections within the Ukhaa Tolgod drainage basin. The stratigraphic sections were measured with a Jacob staff and Brunton compass (fig. 8). The location of each can be seen on maps 3–Map 4Map 56.
Downlap Hill
Figure 9 (top and bottom) depicts the lateral relationships at the site, where Facies E-1 forms the base of the western edge of the hill (map 3). The overlying Facies S contains faint, low-angle bedding planes that dip 5–10 degrees northeast and may represent contacts between individual debris flows that formed the alluvial fan. Above, cross-strata of Facies E-1 fills in a shallow depression that formed the surface of the fan. They dip about 25 degrees to the northeast at a bearing of 55 degrees in relation to magnetic north, and clearly represent the downlap of the dune over the fan surface as it migrated to the northeast.
Figure 8 documents eight sections along the main axis of the drainage basin, from Delta Force/Granger's Hill in the southwest (at left in the illustration) to First Strike in the northeast (at center right). Three lithologic units are especially useful for establishing lithostratigraphic correlations between these sections. First, a distinctive bed containing abundant, small, white, caliche pebbles can be lithostratigraphically correlated from Granger Hill to First Strike. Second, a distinctive interval containing bed(s) of Facies M is consistently found below the white, caliche pebble layer at Ankylosaur Flats, Camel Humps, SW of Xanadu, and First Strike. Finally, a lithologically distinct bed of Facies C containing andesitic clasts caps the sections at Ankylosaur Flats and Camel Humps.
Delta Force/granger Hill
Vertebrate fossils including an articulated oviraptorid skeleton are common in the basal 7 m at Granger Hill (map 3). Sandslides of Facies S, which cap the section at Delta Force (fig. 10), have produced a well-preserved articulated skeleton of Deltatheridium. A pair of intertwined oviraptorid skeletons was excavated at Romeo and Juliet to the northeast (map 3).
Ankylosaur Flats
This stratigraphic section runs from near the local bottom of the Ukhaa Tolgod drainage basin to the top of the bluffs overlooking Ankylosaur Flats (fig. 11, top; map 4). The lateral relationships of strata exposed along the ridge between Ankylosaur Flats and the Sphinx are illustrated in figure 11 (bottom). The strata pictured form the basis for the Ankylosaur Flats and Sphinx sections.
Basal concretionary sheets of E-2 in the southwest correlate laterally with Facies S to the northeast, which crops out below a lens of Facies M. Above, in situ remains of articulated partial ankylosaur skeletons are especially abundant on the flats formed by Facies S, while several in situ articulated, oviraptorid specimens, some preserved in brooding position atop nests of eggs, have been excavated from Facies S just below the unit of Kwp. Numerous small concretions containing well-preserved skulls of mammals and lizards occur as float and cannot be assigned with certainty to this stratigraphic interval. In general, this fossil locality is called Ankylosaur Flats.
On the slope below the middle portion of the ridge, another thin vaguely cross-stratified unit crops out, representing the northeast migration of a small dune across the sandslide sequence. The zone of Kwp marks a period of incipient soil formation at the top of this lower sandslide sequence. Above Kwp, another unit of Facies S contains numerous partially articulated oviraptorid specimens.
Along the southwestern half of the ridge above Kwp, units of Facies S run all the way up to the base of the capping Facies C. However, to the northeast a prominent unit of Facies E-1 is exposed, which forms the bulk of Sphinx Butte and clearly represents the northeast migration of a large dune over the laterally equivalent units of Facies S exposed along the ridge to the southwest.
Sphinx
A butte, called the Sphinx (fig. 12, map 4), juts out from the northern edge of the Ukhaa Tolgod drainage basin. Individual beds of Facies S at the base attain thicknesses of 50 to 75 cm, possibly providing some sense of the thickness of distinct debris flow units that moved down the dunes. Nests of dinosaur eggs and abundant eggshell fragments, along with articulated partial oviraptorid skeletons are common.
Above, cross strata of E-1, equivalent to the interval of Facies S between the 26 and 38 m marks in the Ankylosaur Flats section, are occasionally deformed by concave-up folds of various sizes interpreted to represent tracks of large vertebrates (Loope et al., 1998). Rhizoliths are the only other fossils present.
Facies S caps the Sphinx section and contains calcareous nodules that occasionally yield skulls of mammals and lizards. Although they usually occur as float, they can be assigned to this interval because it represents the top of the section.
Sugar Mountain
Between the Sphinx and Camel Humps lie intermittent exposures of Facies S (fig. 13, map 4). These represent the renewed mass wasting deposits at the base of the dunes represented by Facies E-1 at the Sphinx. Associated remains of Protoceratops are especially abundant, and near the top of the interval, several, closely associated and articulated skeletons of small placental mammals including Ukhaatherium nessovi Novacek et al., 1997 ( = gen. nov., Dashzeveg et al., 1995: 448) have been collected.
Camel Humps Basin
The basal exposures contain beds of Facies S interbedded with interdune deposits of Facies M (fig. 14, map 4), which lithostratigraphically correlates with those near the base of the eastern portion of the Ankylosaur Flats section. These mudstones have yielded two fossil turtles, with preserved shells and other postcrainal remains, as well as a few partial skeletons of Protoceratops. The basal units of M and S (fig. 15) can be traced northeast along the base of the ridge where they abut a unit of E-1. Just above the basal mudstones to the south of Camel Humps on a small ridge jutting out from the base of the buttes, Kwp contains abundant fossil skulls of lizards at Jim's Hot Spot.
The northern ridge of the basin contains many of the same units seen in the southern ridge. The unit of E-1 seen below Camel Humps Buttes can be traced through the small drainage of the basin to where it dips eastward across the northern ridge, lapping down across the basal, fossiliferous unit of Facies S (fig. 16, top). This unit of E-1 laterally correlates with the E-1 unit that forms Sphinx Butte and appears to represent a remobilization of that dune. However, the deposition of the E-1 unit at Camel Humps postdated that at the Sphinx, because the bearings and dips of the cross-strata indicate a northeast migration of the dunes with an intervening period of sandslide development between Sugar Mountain and Camel Humps. This dune appears to have downlapped over the intervening sandslide deposits between the Sphinx and Camel Humps. The period of dune remobilization was not extensive, however, as Facies S comprises most of the laterally equivalent deposits along the northern ridge.
Above, a unit of Facies S is laterally equivalent to that at the top of the Sphinx. Near the middle of this unit at Death Row, numerous associated and articulated oviraptorid specimens, as well as specimens of Protoceratops, an ankylosaur, and a dinosaur nest were collected (fig. 16, bottom). The fossils occur within a very limited interval, suggesting that this level may have represented a distinct topographic surface on the sandslide deposits. Above that interval, occasional lenses of igneous pebbles and isolated cobbles, identical to those in the capping unit of Facies C, appear to document the approach of a coarse-grained alluvial fan from outside the dunefield that eventually overran this portion of the dunefield and deposited the capping conglomerates. The compositional similarities strongly suggest that the capping conglomerate is Cretaceous.
Xanadu Southwest
The basal Facies S has yielded a well-preserved ankylosaur skull. Above, Facies M correlates with basal beds of M on the southern side of Camel Humps Basin. The section is capped by Facies E-2 (fig. 17, map 5).
Xanadu
The section's base, formed by E-2, lies at the bottom of the main drainage for the Ukhaa Tolgod Basin (map 5). But to the north, Facies S forms the base of the section and appears to represent sandslides building out from the laterally equivalent dune sands to the west. Calcareous nodules in Facies S have produced a well-preserved multituberculate skull and abundant eggshell fragments.
Above the white pebble layer (fig. 18), fossils from Facies S at Xanadu include abundant dinosaur nests, an oviraptorid embryo, partially articulated ankylosaur skeletons, Protoceratops skeletons, juvenile dromaeosaur skulls, and skulls of mammals and lizards in nodular calcareous concretions.
First Strike
The basal unit of E-2 (fig. 19A, map 5) probably corresponds laterally to the E-2 at the base of Xanadu. The Facies M forming the base of the butte at First Strike correlates with the basal M at Camel Humps and Xanadu SW. Lizard and mammal skulls from the overlying Facies S are preserved in calcareous, nodular concretions as float, and Kwp occurs near the top of this unit (fig. 19B). A stratigraphically higher unit of Facies S produced numerous fossil skulls of lizards and mammals preserved in nodular calcareous concretions. These included the first fossils discovered in the Ukhaa Tolgod Basin in 1993.
Zofia's Hill, Zofia's Middle, and Zofia's North
Figure 8 (far right in illustration) and map 6 document three stratigraphic sections along the northernmost extent of the Ukhaa Tolgod drainage basin between Zofia's Hill in the southwest (fig. 20A), Zofia's Middle in between (fig. 20B), and Zofia's North in the northeast.
The basal Facies S at Zofia's Hill yielded numerous skulls of mammals and lizards in nodular calcareous concretions.
Six m above the prominent unit of Facies C at Zofia's Middle (fig. 20B) lies a unit of Facies S, containing nodular calcareous concretions occurring as float with fossil skulls of lizards and mammals. This unit correlates with the fossiliferous unit of Facies S at Zofia's Hill.
The thin bed of Facies C at Zofia's North can be laterally traced into the similar unit at the base of the Zophia's Middle section. An overlying unit of Facies S contains calcareous nodules occurring as float that contain fossil skulls of lizards.
At present, no distinctive lithologic units have been identified that can be used to establish lithostratigraphic correlations between the three sections in the Zofia's Hill region and the eight sections to the south. However, measurement and calculation of the strata's local dip can be used to infer the superpositional relationship.
Calculation of Local Dip and Its Implication for the Stratigraphic Position of the Sections at Zofia's Hill
The local dip of the lithostratigraphic units can be estimated in two ways that yield very similar results. First, the bed of conglomerate in the Zofia's Middle section is exposed over a distance of about 100 m in the drainage trending north-south that runs along the western margin of the outcrops at Zofia's Hill. Using the inclinometer on a Brunton compass, the apparent dip of the conglomerate, as seen in cross section within the outcrop, was estimated to be about 2.5 degrees to the south.
This estimate corresponds closely with the dip of the siltstones exposed in the areas around Camel Humps and Xanadu SW sections, as calculated on the basis of a geometric construction using the physiographic map (map 1). Using an altimeter, the elevations of the siltstones were estimated at the base of the southern ridge at the south end of Camel Humps Basin, on the northern flank of Camel Humps Basin, and at Xanadu SW. These siltstones are lithostratigraphically correlated with one another because they are the only siltstones exposed in each section, and they immediately underlie the zone of small, white, caliche pebbles in each section. The gradient observed between the exposures at the northern and southern ends of the Camel Humps Basin (line A–B) was used to estimate the position of the siltstone at the same elevation of the siltstone exposed at Xanadu SW. A line connecting points of equal elevation in the Camel Humps and Xanadu SW areas (line B–C) defines the strike of the siltstone—about 70° NW. The dip of the siltstone, based on calculating the tangent of the angle of dip from the elevation difference between the siltstones on the southern edge of Camel Humps Basin and the elevation along the line of strike divided by the distance between the line of strike and the siltstone on the southern flank of Camel Humps Basin (line A–D), is 2.4° to the SW.
Knowing the strike and dip of the beds in the Ukhaa Tolgod Basin allows one to calculate the projected stratigraphic position of the units at Zofia's Hill in relation to those at Camel Humps. A line of strike is drawn through the conglomerate exposure near Zofia's Hill (line E–F). Line A–F, which is perpendicular to E–F then represents the direction of dip. Knowing the angle of dip (2.4°) and the distance between the line of strike (E–F) and the exposures of siltstone at Camel Humps (line A–F) allows one to calculate the projected elevation of Zofia's conglomerate at Camel Humps. The calculation suggests that the conglomerate in the Zofia's Middle section and the laterally equivalent siltstone in the Zofies North section would fall about 28 m below the mudstone at the southern edge of Camel Humps Basin.
Discussion
Comments on Fossil Preservation
Although none of the fossils at Ukhaa Tolgod were collected from strata that can, using the well-tested criteria of Hunter (1977, 1981), be interpreted as eolian, they commonly do lie adjacent to eolian deposits (see maps 3–6). Can the debris flow hypothesis developed at Ukhaa Tolgod explain the occurrence of fossils at other Gobi localities? Fastovsky et al. (1997) report the presence of abundant vertebrate fossils in cross-stratified sands at Tugrugyin-Shireh. The cross-stratified sands are interpreted to represent the migration of eolian dunes, and we concur. Structureless sandstones are, however, also present at Tugrugyin-Shireh, and the articulated dinosaur skeletons we observed during our reconnaissance in July 1998 lie within these structureless sands. If vertebrate fossils are preserved within (as opposed to adjacent to) cross-stratified eolian sands at Tugrugyin-Shireh as reported by Fastovsky et al. (1997), this contrasts with the lack of vertebrate fossils in cross-stratified eolian sands at Ukhaa Tolgod (Loope et al., 1998, 1999, and this paper) and Bayan Mandahu (Eberth, 1993).
Eberth (1993) describes both cross-stratified and structureless sandstones at Bayan Mandahu. Whereas the structureless sandstones contain vertebrate fossils, the cross-stratified sandstones contain only trace fossils. Eberth interprets both facies to represent products of eolian dune deposition, with structureless sands possibly representing interdune deposits of zibars—low relief dunes composed of grains too coarse to allow the development of a slip face (Kocurek, 1996). We concur with the assignment of the cross-stratified sand to eolian facies. However, we are posing an alternative explanation for the structureless sands at Ukhaa Tolgod, which we feel are better explained as sandslide deposits built up at the base of dunes.
Talbot (1985), in his discussion of Quaternary cycles of dune mobility and stabilization, argued that “dinosaurs in Mongolia were hatching and raising their young in a moist, stabilized dunefield rather than an active sea”. Our observations on the clay-coated sand grains, pedogenic calcite, bioturbation, and lateral relationships between the dune and sandslide facies lead us to agree that this is the case at Ukhaa Tolgod. Although the travel literature of Central Asia and Arabia commonly cites heat, cold, and drought as causes for mass mortality, Finch et al. (1972) found no reports documenting the smothering of animals during desert dust storms and sandstorms. Although they contain abundant signs of life, the cross-stratified eolian facies at Ukhaa Tolgod (E-1 and E-2) are essentially devoid of skeletal remains. If stratal geometry and taphonomic evidence were not considered, the structureless sandstones of Facies S might be interpreted as eolian sediment that accumulated slowly enough for bioturbation to keep pace with deposition. However, it is unlikely that small, delicate mammal and lizard fossils would have been preserved essentially intact after extensive bioturbation. The abundant articulated skeletons are, thus, clear evidence for rapid deposition. The absence of skeletons in the eolian strata that show evidence for rapid emplacement (Facies E-1) is inconsistent with the “sandstorm” hypothesis for fossilization.
Our sandslide interpretation for Facies S provides an alternative to the sandstorm (eolian) hypothesis for in situ burial of articulated skeletons. The high, sparsely vegetated landforms were a sediment reservoir available for episodic gravity flows capable of quickly burying intact, unscavenged skeletons and, possibly, live animals. Heavy rainfall events acted as the triggering mechanism. However, the low permeability of calcite-cemented zones at shallow depth below lee slopes of dunes is the sine qua non of the lethal mass wasting.
Comments on Formational Assignment and Geologic Age
The question arises as to whether the strata at Ukhaa Tolgod are best referred to as extensions of the type Djadokhta or Barun Goyot Formation. A preliminary investigation referred the exposures at Ukhaa Tolgod to the Djadokhta Formation (Dashzeveg et al., 1995).
Such an assignment must be based primarily on lithologic criteria, with fossil content playing a subordinate role, if any (Hedburg, 1976). Gradziński et al. (1977) provided diagnoses for the Djadokhta and Barun Goyot Formations; however, they noted (Gradziński et al., 1977: 307), “No physical contact between the Djadokhta and Barun Goyot formations has yet been discovered”. To our knowledge, that is, unfortunately, still the case today, making assignment of the strata at Ukhaa Tolgod more difficult.
Yet, similar to the classic Djadokhta exposures at Bayn Dzak, the sequence of strata at Ukhaa Tolgod reflects a mixture of facies E-1, E-2, F, and S in the lower part of the section, with prominent beds of Facies C near the top (Gradziński et al., 1969, 1977; Gradziński, 1970; Dashzeveg et al., 2005). Dashzeveg et al. (2005) refer the exposures at Bayn Dzak to the Bayn Dzak Member of the Djadokhta Formation. Such a sequence has yet to be recognized in classic Barun Goyot exposures at Khulsan or Khermeen Tsav (Gradziński and Jerzykiewicz, 1974a, 1974 b; Gradziński et al., 1977). Instead, as those authors state (Gradziński et al., 1977: 287), “Flat-bedded sandstones intercalated with mudstones and/or siltstones become dominant in the uppermost part of the section”. Hence, the sequence of strata at Ukhaa Tolgod conforms more closely to the sequence of the Bayn Dzak Member of the Djadokhta Formation at Bayn Dzak. Given that neither the Bayn Dzak Member nor the Djadokhta Formation has been previously recognized in the Nemegt Basin, one might question whether such a large geographic extension southward from the Ulan Nur Basin can be justified. However, correlative strata of the Djadokhta Formation have already been recognized even further to the south at Bayan Mandahu in Chinese Inner Mongolia (e.g., Jerzykiewicz et al., 1993a, 1993b). Given these points, we refer the strata at Ukhaa Tolgod to the Bayn Dzak Member of the Djadokhta Formation.
Makovicky 2007 developed “cladistic biochronologic analysis”, which uses phylogenetic branching pattern to temporally order paleobiotas. This technique is based on Brook's parsimony analysis (Brooks, 1990) and allows faunas to be temporally ordered if a good fossil record is present, even if no geochronologic evidence is available. Note, however, that while this method orders different faunas it does not indicate absolute age or amount of time between faunas. Makovicky's analysis of Djadoktha localities was based on lizards, dinosaurs, mammals and turtles. He showed that using the CBA method, the Ukhaa Tolgod fauna clustered with localities classically considered to be Barungoyotian in age, younger than other classic Djadoktha localities like Bayn Dzak.
Faunas from the Djadokhta Formation have been assigned a Campanian–Early Maastrichtian age by Lillegraven and McKenna (1986) and Averianov (1997), based primarily on biochronologic interpretations. Using a line of evidence independent of biochronologic interpretations to test these assignments, Dashzeveg et al. (2005) assign a late Campanian age to the strata of the Bayn Dzak Member at Bayn Dzak based on tentative magnetostratigrapic correlations to the Geomagnetic Polarity Timescale (Gradstein et al., 1995). Further, in their analysis of the lizard faunas from the Djadokhta and Barun Goyot Formations, Gao and Norell (2000) state, “lizards provide little evidence for temporally ordering or subdividing Barun Goyot and Djadokhta localities, since 15 of the 19 (78.9%) lizard species that occur at more than one locality are found in both Barun Goyot and Djadokhta localities”. Based on the superpositional relationship between the Tugrugyin Member and Bayn Dzak Member proposed by Dashzeveg et al. (2005), the fauna from Tugrugyin Shireh must be somewhat younger than that from Bayn Dzak, although the extent of this age difference is presently uncertain. Regarding the mammalian faunas from Bayn Dzak, Tugrugyin Shireh, and Ukhaa Tolgod, the extensive variation in form observed in the large sample of multituberculate skulls from Ukhaa Tolgod is now being analyzed systematically. Those analytical results could well affect the current taxonomic identifications for multituberculates from all three localities. Consequently, pending further evidence and analysis, the vertebrate fauna at Ukhaa Tolgod (table 1) is thought here to reflect a Campanian age.
Table 1
Vertebrate Faunal List for Ukhaa Tolgod
Conclusions
The deposits exposed in the Ukhaa Tolgod Basin represent a mosaic of previously described cross-stratified dune units, siltstone interdune deposits, and conglomeratic beds washed in from the basin margin, mixed with structureless sands interpreted here to represent a newly recognized facies of sandslides deposited at the base of the dunes (S). Vertebrates traversed the dunefield when it was active, as evidenced by the concave-up depressions interpreted to represent their tracks. However, their fossilized skeletal remains were preserved exclusively in the sandslide facies (including admixed fluvial deposits). Their preservation seems to be associated with the sandy mass flows that formed the fans during periods when dunes were stabilized and after pedogenic calcite had accumulated under dune slopes. Rainfall saturated the upper portion of the dunes and triggered the debris flows. The dunes were sparsely vegetated, as evidenced by the occasional zones of rhizoliths preserved in the cross-stratified sands. The distinctly cross-stratified sands (E-1) accumulated during more arid periods when dunes were mobile; burrowing and calcite accumulation (Facies E-2) took place when dunes were stabilized. Ephemeral small lakes or ponds occasionally occupied the interdune corridors and received silty muds and sands (Facies M). Both before and after the dunefield formed, coarse clasts derived from nearby topographic highs washed into the area of the dunefield from the basin margin.
Based on superpositional similarities of these facies at Ukhaa Tolgod and Bayn Dzak, the exposures at Ukhaa Tolgod are assigned to the Bayn Dzak Member of the Djadokhta Formation. At both localities, facies E-1, E-2, S, and M dominate the lower portions of the sections, with prominent beds of Facies C exposed near the top. Exposures of the Barun Goyot Formation at Khulsan differ substantially, with units of flat-bedded sandstone intercalated with beds of Facies M dominating the upper portion of the sequence (Gradziński et al., 1977).
The vertebrate fauna from Ukhaa Tolgod is tentatively assigned a Campanian age, based on recent biochronologic and magnetostratigraphic analyses (Lillegraven and McKenna, 1986; Averianov, 1997; Gao and Norell, 2000; Dashzeveg et al., 2005).
Acknowledgments
The authors wish to thank the Mongolian Academy of Sciences, as well as the field crews from the American Museum of Natural History. We also wish to thank Zofia Kielan-Jaworowska, Jason Lillegraven, David Eberth, Dale Russell, and one anonymous reviewer for their extensive reviews of this manuscript, without implying that they concur with all the statements and conclusions presented herein. Illustrations for both this paper and Dashzeveg et al., 2005, were compiled and drafted by Frank Ippolito, and assistance in compiling the faunal lists was generously provided by James Clark and Gao Keqin. Financial support for this study was provided by the Frick Endowment to the Division of Paleontology at the American Museum of Natural History, the InfoQuest Foundation, the Ann and Gordon Getty Foundation, and the Charlotte and Walter Kohler Charitable Trust.