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Type: Article
Published: 2019-05-17
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Recommended abbreviations for the names of genera of the phylum Tardigrada

Department of Biology, Unity College, Unity, ME 04988 U.S.A.
Department of Biology and Chemistry, Baker University, Baldwin City, KS 66006, U.S.A.
Department of Animal Taxonomy and Ecology, Adam. Mickiewicz University, Poznań, Umultowska 89, 61-614 Poznań, Poland.
Tardigrada tardigrades nomenclature abbreviations genera history ICZN

Abstract

When listing the tardigrade species found in a study, the general practice is to sort the list alphabetically or taxonomically by genus. The first time a genus name is used it is spelled out and then it is abbreviated with the first letter of the genus followed by a full stop (period) until a different genus is encountered. This practice works for both a paragraph and a table listing and has been used by tardigrade researchers as far back as Schultze (1834). In the last monograph, Ramazzotti & Maucci (1983) presented a listing of the 52 generic abbreviations they used. We present an updated list of unambiguous abbreviations consistent with the International Code of Zoological Nomenclature (ICZN) for the now 137 recognized tardigrade genera.

 

References

  1. Bertolani, R. (2003) Thulinius, new generic name substituting for Thulinia Bertolani, 1981 (Tardigrada, Eutardigrada). Zootaxa, 314, 1–4.

    Bertolani, R. & Biserov, V.I. (1996) Leg and claw adaptations in soil tardigrades, with erection of two new genera of Eutardigrada, Macrobiotidae: Pseudohexapodibius and Xerobiotus. Invertebrate Biology, 115 (4), 299–304.

    https://doi.org/10.2307/3227019

    Bertolani, R. & Pilato, G. (1988) Struttura delle unghie nei Macrobiotidae e descrizione di Murrayon n. gen. (Eutardigrada). Animalia, 15 (1–3), 17–24.

    Bertolani, R., Guidetti, R., Marchioro, T., Altiero, T., Rebecchi, L. & Cesari, M. (2014) Phylogeny of Eutardigrada: New molecular data and their morphological support lead to the identification of new evolutionary lineages. Molecular Phylogenetics and Evolution, 76, 110–126.

    https://doi.org/10.1016/j.ympev.2014.03.006

    Binda, M.G. (1978) Risistemazione di alcuni Tardigradi con l'istituzione di un nuovo genere di Oreellidae e della nuova famiglia Archechiniscidae. Animalia, 5 (1–3), 307–314.

    Binda, M.G. & Pilato, G. (1986) Ramazzottius, nuovo genere di eutardigrado (Hypsibiidae). Animalia, 13 (1–3), 159–166.

    Biserov, V.I. (1992) A new genus and three new species of tardigrades (Tardigrada: Eutardigrada) from the USSR. Bollettino di Zoologia, 59 (1), 95–103.
    https://doi.org/10.1080/11250009209386654

    Bussau, C. (1992) New deep-sea Tardigrada (Arthrotardigrada, Halechiniscidae) from a manganese nodule area of the eastern South Pacific. Zoologica Scripta, 21 (1), 79–91.
    https://doi.org/10.1111/j.1463-6409.1992.tb00311.x

    Cantacuzène, M.A. (1951) Tardigrade marin nouveau, commensal de Limnoria lignorum (Rathke). Comptes Rendus de l’Académie des Sciences, 232, 1699–1700.

    Claxton, S.K. (1999) Milnesioides exsertum gen. n. sp. n., a new tardigrade from Australia (Tardigrada: Milnesiidae). Zoologischer Anzeiger, 238 (3–4), 183–190.

    Cooper, K.W. (1964) The First Fossil Tardigrade: Beorn leggi Cooper, from Cretaceous amber. Psyche, 71, 41–48.

    https://doi.org/10.1155/1964/48418.

    Cuénot, L. (1892) Commensaux et parasites des Echinodermes (21ème note). Revue de Biologie du Nord de la France, 5, 1–22.

    Cuénot, L. (1926) Description d'un Tardigrade Nouveau de la Faune Francaise. Comptes rendus de l'Académie des Science, Paris, 182, 744.

    D'Addabbo Gallo, M., De Zio Grimaldi, S., Morone De Lucia, M.R. & Troccoli, A. (1992) Halechiniscidae and Echiniscoididae from the Western Mediterranean Sea. (Tardigrada: Heterotardigrada). Cahiers de Biologie Marine, 33, 299–318.
    https://doi.org/10.21411/CBM.A.D1AD0BC1

    Dantas-Torres, F. (2008) Towards the standardization of the abbreviations of genus names of ticks (Acari:Parasitiformes:Ixodida). Parasitologia Veterinária, 154 (1–2), 68–76

    Dastych, H. (1983) Apodibius confusus gen. n. sp. n., a new water-bear from Poland (Tardigrada). Bulletin of the Polish Academy of Sciences, 31 (1–2), 41–46.

    Dastych, H. (1992) Paradiphascon manningi gen. n. sp. n., a new water-bear from South Africa, with the erecting of a new subfamily Diphasconinae (Tardigrada). Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, Hamburg, 89, 125–139.

    Dastych, H. (1993) A new genus and four new species of semiterrestrial water-bears from South Africa (Tardigrada). Mitteilungen aus den Hamburgischen Zoologischen Museum und Institut, Hamburg, 90, 175–186.

    Dastych, H. (2009) Thalerius konradi gen. nov., sp. nov., a new tardigrade from the periglacial area of the Ötztal Alps, Austria (Tardigrada). Contributions to Natural History, 12, 391–402.

    Dastych, H. (2011) Bergtrollus dzimbowski gen. n., sp. n., a remarkable new tardigrade genus and species from the nival zone of the Lyngen Alps, Norway (Tardigrada: Milnesiidae). Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 15, 335–359.

    de Barros, R. (1939) Itaquascon umbellinae gen. nov. spec. nov. (Tardigrada, Macrobiotidae). Zoologischer Anzeiger, 128 (3–4), 106–109.

    de Zio Grimaldi, S., D'Addabbo Gallo, M. & Morone De Lucia, M.R. (1992) Neoarctus primigenius n.g., n.sp., a new Stygarctidae of the Tyrrhenian Sea (Tardigrada, Arthrotardigrada). Bollettino di Zoologia, 59, 309–313.
    https://doi.org/10.1080/11250009209386687

    Degma, P. & Guidetti, R. (2007) Notes to the current checklist of Tardigrada. Zootaxa, 1579, 41–53.

    Degma, P., Bertolani, R. & Guidetti, R. (2009–2018) Actual checklist of Tardigrada species. 34th Edition. 30 June 2018. Available from: http://www.tardigrada.modena.unimo.it/miscellanea/Actual%20checklist%20of%20Tardigrada.pdf (accessed 15 January 2019)

    Delamare Deboutteville, C. & Renaud-Mornant, J. (1965) Un remarquable genre de Tardigrades des sables coralliens de Nouvelle-Calédonie. Bulletin du Muséum National D'Histoire Naturelle, Section A Zoologie Biologie et Ecologie Animales, 260 (2), 2581–2583.

    Doyère, L.M.F. (1840) Mémoire sur les Tardigrades. I. Annales des Sciences Naturelles, Paris, Series 2, 14, 269–362.

    du Bois-Reymond Marcus, E. (1944) Sobre tardigrados Brasileiros. Communicaciones Zoologicas del Museo de Historia Natural de Montevideo, 1 (13), 1–19.

    du Bois-Reymond Marcus, E. (1952) On South American Malacopoda. Boletim da Faculdade de Filosofia, Ciências e Letras da Universidade de São Paulo, Serie Zoologia, 17, 189–209.

    Ehrenberg, C.G. (1848) Fortgesetze Beobachtungen über jetzt herreschende atmosphãrische mikroscopische, etc. mit Nachtrag und Novarum Specierum Diagnosis. Bericht über die zur Bekanntmachung geeigneten Verhanlungen del Koniglichen Preussischen Akademie der Wissenschaften zu Berlin, 13, 370–381.

    Ehrenberg, C.G. (1859) Beitrag zur Bestimmung des stationären mikroscopischen Lebens in bis 20,000 Fuss Alpenhöhe. Abhandlungen der Königlichen Akademie der Wissenschaften zu Berlin, 1859, 429–456.

    Fujimoto, S. (2015) Quisarctus yasumurai gen. et sp. nov. (Arthrotardigrada: Halechiniscidae) from a submarine cave, off Iejima, Ryukyu Islands, Japan. Zootaxa, 3948 (1), 145–150.

    https://doi.org/10.11646/zootaxa.3948.1.10

    Fujimoto, S. & Yamasaki, H. (2017) A new genus and species of Renaudarctidae (Heterotardigrada: Arthrotardigrada) from Ryukyu Archipelago, Japan. Marine Biology Research, 13 (3), 288–299.

    https://doi.org/10.1080/17451000.2016.1257809

    Gąsiorek, G., Stec, D., Morek, W. & Michalczyk, Ł. (2019) Deceptive conservatism of claws: distinct phyletic lineages concealed within Isohypsibioidea (Eutardigrada) revealed by molecular and morphological evidence. Contributions to Zoology, 1–55. [published online]
    https://doi.org/10.1163/18759866-20191350

    Gąsiorek, P., Morek, W., Stec, D. & Michalczyk, Ł. (2019) Untangling the Echiniscus Gordian knot: paraphyly of the “arctomys group” (Heterotardigrada: Echiniscidae). Cladistics. [published online]
    https://doi.org/10.1111/cla.12377

    Gąsiorek, P., Suzuki, A.C., Kristensen, R.M., Lachowska-Cierlik, D. & Michalczyk, Ł. (2018) Untangling the Echiniscus Gordian knot: Stellariscus gen. nov. (Heterotardigrada: Echiniscidae) from Far East Asia. Invertebrate Systematics, 32, 1234–1247.

    Grimaldi de Zio, S., D’Addabbo Gallo, M. & Morone De Lucia, R.M. (1982) Neostygarctus acanthophorus, n. gen. n. sp., nuovo tardigrado marino del Mediterraneo. Cahiers de Biologie Marine, 23, 319–323.

    https://doi.org/10.21411/CBM.A.66544B89

    Gross, V., Miller, W.R. & Hochberg, R. (2014) A new tardigrade, Mutaparadoxipus duodigifinis gen. nov., sp. nov. (Heterotardigrada: Arthrotardigrada) from the southeastern United States. Zootaxa, 3835 (2), 263–272.

    https://doi.org/10.11646/zootaxa.3835.2.6

    Guidetti, R. & Bertolani, R. (2005) Tardigrade taxonomy: an updated check list of the taxa and a list of characters for their identification. Zootaxa, 845 (1), 1–46.
    https://doi.org/10.11646/zootaxa.845.1.1

    Guidetti, R. & Pilato, G. (2003) Revision of the genus Pseudodiphascon (Tardigrada, Macrobiotidae), with the erection of three new genera. Journal of Natural History, 37 (14), 1679–1690.

    https://doi.org/10.1080/00222930210126659

    Guidetti, G., Rebecchi, L., Bertolani, R., Jönsson, K.I., Kristensen, R.M. & Cesari, M. (2016) Morphological and molecular analyses on Richtersius (Eutardigrada) diversity reveal its new systematic position and lead to the establishment of a new genus and a new family within Macrobiotoidea. The Linnean Society of London, Zoological Journal of the Linnean Society, 178, 834–845.

    https://doi.org/10.1111/zoj.12428

    Guidetti, R., Schill, R.O., Bertolani, R., Dandekar, T. & Wolf, M. (2009) New molecular data for tardigrade phylogeny, with the erection of Paramacrobiotus gen. nov. Journal of Zoological Systematics and Evolutionary Research, 47 (4), 315–321. https://doi.org/10.1111/j.1439-0469.2009.00526.x

    Guil, N., Rodrigo, E. & Machordom, A. (2014) Soil tardigrade biodiversity with the description of a new eutardigrade genus and its phylogenetic position. Systematics and Biodiversity, 13 (3), 234–256.

    https://doi.org/10.1080/14772000.2014.986554

    Hansen, J., Kristensen, R. & Jørgensen, A. (2012) The armored marine tardigrades (Arthrotardigrada, Tardigrada). Scientia Danica, Series B, Biologica, 2, 1–91.

    Horning, D.S.J., Schuster, R.O. & Grigarick, A.A. (1978) Tardigrada of New Zealand. New Zealand Journal of Zoology, 5, 185–280.
    https://doi.org/10.1080/03014223.1978.10428316

    ICZN (1999) International Code of Zoological Nomenclature. 4th Edition. The International Trust for Zoological Nomenclature, London, 306 pp.

    https://doi.org/10.5962/bhl.title.50608

    Kaczmarek, Ł. & Michalczyk, Ł. (2006) Redescription of Macrobiotus tridigitus Schuster, 1983 and the erection of a new genus of Tardigrada (Eutardigrada, Macrobiotidae). Journal of Natural History, 40 (19–20), 1223–1229.

    https://doi.org/10.1080/00222930600844443

    Kristensen, R.M. (1982a) New aberrant eutardigrades from homothermic springs on Disko Island, West Greenland. In: Proceedings of the Third International Symposium on the Tardigrada, Johnson City, Tennessee, 1982, pp. 203–220.

    Kristensen, R.M. (1982b) The first record of cyclomorphosis in Tardigrada based on a new genus and species from Arctic meiobenthos. Zeitschrift für Zoologische Systematik und Evolutionsforschung, 20 (4), 249–270.

    https://doi.org/10.1111/j.1439-0469.1983.tb00552.x

    Kristensen, R.M. (1984) On the biology of Wingstrandarctus corallinus nov. gen. et spec., with notes on the symbiotic bacteria in the subfamily Florarctinae (Athrotardigrada). Videnskabelige Meddelelser Dansk naturhistorisk Forening, 145, 201–218.

    Kristensen, R.M. (1987) Generic revision of the Echiniscidae (Heterotardigrada), with a discussion of the origin of the family. In: Bertolani, R. (Ed.), Biology of Tardigrades. Mucchi, Modena, pp. 261–335.

    Kristensen, R.M. & Higgins, R.P. (1984) A new family of Arthrotardigrada (Tardigrada: Heterotardigrada) from the Atlantic coast of Florida, USA. Transactions of the American Microscopical Society, 103 (3), 295–311.

    https://doi.org/10.2307/3226191

    Kristensen, R.M. & Higgins, R.P. (1989) Marine Tardigrada from the Southeastern United States coastal waters. 1. Paradoxipus orzeliscoides n. gen., n. sp. (Arthrotardigrada: Halechiniscidae). Transactions of the American Microscopical Society, 108 (3), 262–282.

    https://doi.org/10.2307/3226344

    Kristensen, R.M. & Renaud-Mornant. J. (1983) Existence d'Arthrotardigrades semi-benthiques de genres nouveaux de la sousfamille des Styraconyxinae subfam. nov. Cahiers de Biologie Marine, 24 (3), 337–353.

    https://doi.org/10.21411/CBM.A.C9AB8A14

    Marcondes, C.B. (2017) A proposal for the use of standardized abbreviations for the genera of triatomine bugs (Reduviidae: Triatominae) across the World. Revista da Sociedade Brasileira de Medicina Tropical, 50 (6), 877–878

    https://doi.org/10.1590/0037-8682-0324-2017

    Marcus, E. (1936) Tardigrada. In: deGruyter, W. (Ed.), Das Tierreich, 66, pp. 1–340. [Berlin und Leipzig]

    Maucci, W. & Ramazzotti, G. (1981a) Adorybiotus gen. nov.: nuova posizione sistematica per Macrobiotus granulatus Richters, 1903 e per Macrobiotus coronifer Richters, 1903 (Tardigrada, Macrobiotidae). Memorie dell’Istituto Italiano di Idrobiologia dott Marco de Marchi, 39, 153–159.

    Maucci, W. & Ramazzotti, G. (1981b) Cornechiniscus gen. nov.: nuova posizione sistematica per i cosiddetti « Pseudechiniscus gruppo cornutus » con descrizionedi una nuova specie (Tardigrada, Echiniscidae). Memorie dell’Istituto Italiano di Idrobiologia dott Marco de Marchi, 39, 147–151.

    May, R.M. (1948) Nouveau genre et espèce de tardigrade du Mexique: Haplomacrobiotus hermosillensis. Bulletin de la Société Zoologique de France, 73, 95–97.

    McKirdy, D.J., Schmidt, P. & McGinty-Bayly, M. (1976) Interstitielle Fauna von Galapagos. XVI. Tardigrada. Mikrofauna des Meeresbodens, Mainz, 58, 409–449.

    Møbjerg, N., Kristensen, R.M. & Jørgensen, A. (2016) Data from new taxa infer Isoechiniscoides gen. nov. and increase the phylogenetic and evolutionary understanding of echiniscoided tardigrades (Echiniscoidea: Tardigrada). Zoological Journal of the Linnean Society, 178, 804–818.

    https://doi.org/10.1111/zoj.12500

    Murray, J. (1905) Tardigrada of the Forth Valley. Annals of Scottish Natural History, 1905, 160–164.

    Murray, J. (1910) Part V Tardigrada. Reports on the Scientific Investigations, 1, 83–187.

    Özdikmen, H. (2008) Bertolanius nom. nov., a replacement name for the genus Amphibolus Bertolani, 1981 (Tardigrada: Parachela) with type species designation. Munis Entomology & Zoology, 3 (1), 330–332.

    Pilato, G. (1969a) Schema per una nuova sistemazione delle famiglie e dei generi degli Eutardigrada. Bollettino delle sedute dell'Accademia Gioenia di Scienze Naturali in Catania, Serie IV, 10 (2), 181–193.

    Pilato, G. (1969b) Su un interessante Tardigrado esapodo delle dune costiere siciliane: Hexapodibius micronyx n. gen. n. sp. Bollettino delle sedute dell'Accademia Gioenia di Scienze Naturali in Catania, Serie IV, 9 (9), 619–622.

    Pilato, G. (1971) Necopinatum mirabile n. gen. n. sp., interessantissimo Eutardigrado incertae sedis. Bollettino delle sedute dell'Accademia Gioenia di Scienze Naturali, Catania, Serie IV, X (10), 861–867.

    Pilato, G. (1987) Revision of the genus Diphascon Plate, 1889, with remarks on the subfamily Itaquasconinae (Eutardigrada, Hypsibiidae). In: Bertolani, R. (Ed.), Biology of Tardigrades. Selected Symposia and Monographs. Vol. 1. U.Z.I., Mucchi, Modena, pp. 337–357.

    Pilato, G. (1992) Mixibius, nuovo genere di Hypsibiidae (Eutardigrada). Animalia, 19, 121–125.

    Pilato, G. (1997) Astatumen, a new genus of the Eutardigrada (Hypsibidae, Itaquasconinae). Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 12, 205–208.

    Pilato, G. (1998) Microhypsibiidae, new family of eutardigrades, and description of the new genus Fractonotus. Spixiana, 21 (2), 129–134.

    Pilato, G. (2009) Bindius triquetrus gen. nov. sp. nov. (Eutardigrada, Hypsibiidae) from Sicily (Italy). Zootaxa, 2058, 62–68.

    Pilato, G. (2013) The taxonomic value of the structures for the insertion of the stylet muscles in the Eutardigrada, and description of a new genus. Zootaxa, 3721 (4), 365–378.
    https://doi.org/10.11646/zootaxa.3721.4.4

    Pilato, G. & Beasley, C.W. (1987) Haplohexapodibius seductor n. gen. n. sp. (Eutardigrada, Calohypsibiidae) with remarks on the systematic position of the new genus. Animalia, 14 (1–3), 65–71.

    Pilato, G. & Binda, M.G. (1987) Parascon schusteri n. gen. n. sp. (Eutardigrada Hypsibiidae, Itaquasconinae). Animalia, 14 (1/3), 91–97.

    Pilato, G. & Binda, M.G. (1989) Richtersius, nuovo nome generico in sostituzione di Richtersia Pilato and Binda, 1987 (Eutardigrada). Animalia, 16, 147–148.

    Pilato, G. & Binda, M.G. (1990) Tardigradi dell'Antartide. I. Ramajendas, nuovo genere di eutardigrado. Nuova posizione sistematica di Hypsibius renaudi Ramazzotti, 1972, e descrizione di Ramajendas frigidus n. sp. Animalia, 17, 61–71.

    Pilato, G. & Binda, M.G. (1997) Acutuncus, a new genus of Hypsibiidae (Eutardigrada). Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 12, 159–162.

    Pilato, G. & Catanzaro, R. (1988) Macroversum mirum n. gen. n. sp. nuovo eutardigrado (Macrobiotidae) dei Monti Nebrodi (Sicilia). Animalia, 15, 175–180.

    Pilato, G. & Lisi, O. (2011) Tenuibiotus, a new genus of Macrobiotidae (Eutardigrada). Zootaxa, 2761, 34–40.

    Pilato, G., Binda, M.G. & Lisi, O. (2004) Famelobiotus salici, n. gen. n. sp., a new eutardigrade from Borneo. New Zealand Journal of Zoology, 31, 57–60.
    https://doi.org/10.1080/03014223.2004.9518359

    Pilato, G., Guidetti, R., Rebecchi, L., Lisi, O., Hansen, J. & Bertolani, R. (2006) Geonemy, ecology, reproductive biology and morphology of the tardigrade Hypsibius zetlandicus (Eutardigrada: Hypsibiidae) with erection of Borealibius gen. n. Polar Biology, 29, 595–603.

    https://doi.org/10.1007/s00300-005-0094-1

    Plate, L. (1888) Beiträge zur Naturgeschichte der Tardigraden. Zoologische Jahrbücher. Abteilung für Anatomie und Ontogenie der Tiere, 3, 487–550.

    Pollock, L.W. (1975) Observations on marine Heterotardigrada including a new genus from the western Atlantic Ocean. Cahiers de Biologie Marine, 16, 121–132.
    https://doi.org/10.21411/CBM.A.71EAFF65

    Pollock, L.W. (1979) Angursa bicuspis n. g., n. sp., a marine arthrotardigrade from the western North Atlantic. Transactions of the American Microscopical Society, 98 (4), 558–562.
    https://doi.org/10.2307/3225907

    Pollock, L.W. (1995) New marine tardigrades from Hawaiian beach sand and phylogeny of the family Halechiniscidae. Invertebrate Biology, 114 (3), 220–235.
    https://doi.org/10.2307/3226877

    Rahm, G. (1937) A new ordo of tardigrades from the hot springs of Japan (Furu-yu section, Unzen). Annotationes Zoologicae Japonensis, 16 (4), 345–352.

    Ramazzotti, G. (1962) Il Phylum Tardigrada. Memorie dell’Istituto Italiano di Idrobiologia, Pallanza, XVI, 1–595.

    Ramazzotti, G. (1965) Il Phylum Tardigrada (1° Supplemento). Memorie dell'Istituto Italiano di Idrobiologia, 19, 101–212.

    Ramazzotti, G. & Maucci, W. (1983) Il Phylum Tardigrada—III edizione. Memorie dell’Istituto Italiano di Idrobiologia, Pallanza, 41, 1–1012.

    Reinert, J.F. (2009) List of abbreviations for currently valid generic-level taxa in family Culicidae (Diptera). European Mosquito Bulletin, 27, 68–76.

    Renaud-Debyser, J. (1959) Études sur la faune interstitielle des Îles Bahamas. III: Tardigrades. Vie et Milieu, 10 (3), 296–302.

    Renaud-Debyser, J. (1965) Parastygarctus higginsi n. g., n. sp. tardigrade marin interstitiel de Madagascar. Comptes rendus de l'Académie des Science, Paris, 260, 955–957.

    Renaud-Mornant, J. (1974) Une nouvelle famille de Tardigrades marins abyssaux: les Coronarctidae fam. nov. (Heterotardigrada). Comptes Rendus de l'Académie des Sciences, Séries D, 278, 3087–3090.

    Renaud-Mornant, J. (1975) Deep-sea Tardigrada from 'Meteor' Indian Ocean Expedition. 'Meteor' Forsch.-Ergebnisse, Séries D, 21, 54–61.

    Renaud-Mornant, J. (1979) Tardigrades marins de Madagascar II. Stygarctidae et Oreellidae. III. Considérations écologiques générales. Bulletin du Muséum National D'Histoire Naturelle, Section A Zoologie Biologie et Ecologie Animales, Séries 4, 8, 339–351.

    Renaud-Mornant, J. (1981) Raiarctus colurus n. g., n. sp., et R. aureolatus n. sp., Tardigrades (Arthrotardigrada) marins nouveaux de sédiments calcaires. Bulletin du Muséum National D'Histoire Naturelle, Section A Zoologie Biologie et Ecologie Animales, 3 (1/2), 515–522.

    Renaud-Mornant, J. (1982) Species diversity in marine Tardigrada. In: Nelson, D. (Ed.), Proceedings of the Third International Symposium on the Tardigrada, East Tennessee State University Press, Johnson City, Tennessee, pp. 149–177.

    Renaud-Mornant, J. (1983) Tardigrades abyssaux nouveaux de la sous-famille des Euclavarctinae n. subfam. (Arthrotardigrada, Halechiniscidae). Bulletin du Muséum National D'Histoire Naturelle, Section A Zoologie Biologie et Ecologie Animales, 5, 201–219.

    Renaud-Mornant, J. (1984) Halechiniscidae (Heterotardigrada) de la campagne Benthedi, canal du Mozambique. Bulletin du Muséum National D'Histoire Naturelle, Section A Zoologie Biologie et Ecologie Animales, Séries 4, 6, 67–88.

    Renaud-Mornant, J. (1987) Halechiniscidae nouveaux de sables coralliens tropicaux (Tardigrada, Arthrotardigrada). Bulletin du Muséum National D'Histoire Naturelle, Section A Zoologie Biologie et Ecologie Animales, Séries 9, 2, 353–373.

    Renaud-Mornant, J. (1989) Opydorscus, un nouveau genre d'Orzeliscinae et sa signification phylogénique (Tardigrada, Arthrotardigrada). Bulletin du Muséum National D'Histoire Naturelle, Section A Zoologie Biologie et Ecologie Animales, Séries 4, 11, 763–771.

    Richters, F. (1908) Marine Tardigraden. Zoologischer Anzeiger, 33, 77–85.

    Richters, F. (1909) Tardigraden-Studien. Bericht der Senckenbergzischen Naturforschenden Gesellschaft in Frankfurt am Main, 1–2, 28–49.

    Rubal, M., Veiga, P., Fontoura, P. & Sousa-Pinto, I. (2013) A new intertidal arthrotardigrade, Prostygarctus aculeatus gen. nov., sp. nov. (Tardigrada: Heterotardigrada) from the north of Portugal (Atlantic ocean). Journal of Limnology, 72 (Supplement 1), 8–14.
    https://doi.org/10.4081/jlimnol.2013.s1.e2

    Schulte, R. & Miller, W.R. (2011) Tardigrades of North America: the elevation of a new genus from California. Transactions of the Kansas Academy of Science, 114 (1–2), 177.

    Schultze, C.A.S. (1834) Macrobiotus hufelandii, animal e crustaceorum classe novum, reviviscendi post diuturnam asphyxiam et ariditatem potens. Apud Carolum Curths, Berolini, 7 pp.

    Schultze, C.A.S. (1840) Echiniscus bellermanni, animal crustaceum, Macrobiotus hufelandii affine. Apud G. Reimer, Berolini, 7 pp.

    Schulz, E. (1935) Actinarctus doryphorus nov. gen. nov. spec., ein merkwürdiger Tardigrad aus der Nordsee. Zoologischer Anzeiger, 111, 285–288.

    Schulz, E. (1951) Über Stygarctus bradypus n.g., n.sp., einen Tardigraden aus dem Küstengrundwasser und seine phylogenetische Bedeutung. Kieler Meeresforschung, 8, 86–97.

    Schulz, E. (1953) Eine neue Tardigraden-Gattung von der pazifischen Küste. Zoologischer Anzeiger, 151, 306–310.

    Schuster, R.O., Nelson, D.R., Grigarick, A.A. & Christenberry, D. (1980) Systematic criteria of the Eutardigrada. Transactions of the American Microscopical Society, 99 (3), 284–303.
    https://doi.org/10.2307/3226004

    Steiner, G. (1926) Bathyechiniscus tetronyx n.g. n. sp. Ein neuer mariner Tardigrade. Deutschen Südpolar-Expedition, 190103, 18, 1–10.

    Thulin, G. (1911) Beitrag zur Kenntnis der Tardigradenfauna Schwedens. Arkiv för Zoologi, Stockholm, 7, 1–60.

    Thulin, G. (1928) Über die phylogenie und das System der Tardigraden. Hereditas Genetiskt Arkiv, 11, 207–266.

    https://doi.org/10.1111/j.1601-5223.1928.tb02488.x

    Thulin, G. (1942) Ein neuer mariner Tardigrad. Meddelanden från Göteborgs Musei Zoologiska Avdelning, 99, 1–10.

    Trygvadóttir, B.V. & Kristensen, R.M. (2011) Eohypsibiidae (Eutardigrada, Tardigrada) from the Faroe Islands with the description of a new genus containing three new species. Zootaxa, 2886, 39–62.

    Vecchi, M., Cesari, M., Bertolani, R., Jönsson, K.I., Rebecchi, L. & Guidetti, R. (2016) Integrative systematic studies on tardigrades from Antarctica identify new genera and new species within Macrobiotoidea and Echiniscoidea. Invertebrate Systematics, 30 (4), 303–322. Available from: http://www.bioone.org/doi/full/10.1071/IS15033 (Accessed 16 May 2019)

    Vicente, F., Fontoura, P., Cesari, M., Rebecchi, L., Guidetti, R., Serrano, A. & Bertolani, R. (2013) Integrative taxonomy allows the identification of synonymous species and the erection of a new genus of Echiniscidae (Tardigrada, Heterotardigrada). Zootaxa, 3613 (6), 557–572.

    https://doi.org/10.11646/zootaxa.3613.6.3

    Villora-Moreno, S. (1996) A New Genus and Species of the Deep-sea Family Coronarctidae (Tardigrada) from a sub-marine cave with a deep-sea like condition. Sarsia, 81 (4), 275–283.

    https://doi.org/10.1080/00364827.1996.10413625

    Zawierucha, K., Stec, D., Lachowska-Cierlik, D., Takeuchi, N., Li, Z. & Michalczyk, Ł. (2018) High mitochondrial diversity in a new water bear species (Tardigrada: Eutardigrada) from mountain glaciers in Central Asia, with the erection of a new genus Cryoconicus. Annales Zoologici, 68 (1), 179–201.

    https://doi.org/10.3161/00034541ANZ2018.68.1.007