Elsevier

Geobios

Volume 54, June 2019, Pages 13-36
Geobios

Research paper
Ammonoids and nautiloids from the earliest Spathian Paris Biota and other early Spathian localities in southeastern Idaho, USA

https://doi.org/10.1016/j.geobios.2019.04.007Get rights and content

Abstract

Intensive sampling of three earliest Spathian sites represented by the Lower Shale unit and coeval beds within the Bear Lake vicinity and neighboring areas, southeastern Idaho, yielded several new ammonoid and nautiloid assemblages. These new occurrences overall indicate that the lower boundary of the Tirolites beds, classically used as a regional marker for the base of the early Spathian, and therefore the regional Smithian/Spathian boundary, must be shifted downward into the Lower Shale unit and coeval beds. Regarding ammonoids, one new genus (Caribouceras) and two new species (Caribouceras slugense and Albanites americanus) are described. In addition, the regional temporal distribution of Bajarunia, Tirolites, Columbites, and Coscaites is refined, based on a fourth sampled site containing a newly reported occurrence of the early Spathian Columbites fauna in coeval beds of the Middle Shale unit. As a complement to ammonoids, changes observed in nautiloid dominance are also shown to facilitate correlation with high-latitude basins such as Siberia during this short time interval, and they also highlight the major successive environmental fluctuations that took place during the late Smithian–early Spathian transition.

Introduction

Pioneering contributions regarding our knowledge of Early Triassic ammonoids from the northern part of the western USA basin were accomplished by White (1879), Hyatt and Smith (1905) and Smith (1932). They were the first to report and illustrate various taxa from the Smithian–Spathian interval in the Bear Lake area of southeastern Idaho (Guex et al., 2010, Jenks et al., 2015; Fig. 1). Smith (1932) also proposed the first regional Early Triassic biozonation based on these organisms by using their successive occurrences observed in several localities within this area. In particular, he recognized the iconic middle and late Smithian Meekoceras and Anasibirites beds, and the early Spathian Tirolites and Columbites beds. Kummel, 1943, Kummel, 1954, Kummel, 1969 continued the exploration of this area and completed the biozonation for the middle Spathian. Kummel, 1954, Kummel, 1957 also documented the major Lower Triassic lithologic units found in the Bear Lake area, provided ammonoid zonal-based ages for each unit and suggested correlation of these units between studied sections (Fig. 2). Additionally, Kummel (1953) reviewed the rare Early Triassic coiled nautiloid occurrences known for this area and noted that a large majority came from the early Spathian Columbites beds. Recently, Guex et al., 2005a, Guex et al., 2005b, Guex et al., 2010 and Jenks et al. (2013) significantly refined these previous works and provided an ammonoid zonation for the Spathian, with particular emphasis on the early Spathian, achieving a much higher resolution than elsewhere in the world (see Jenks et al., 2015 for a review of global Early Triassic correlations).

Here we focus on ammonoids and nautiloids recently found in the earliest Spathian of the Bear Lake area, which yields specimens of both clades (Brayard et al., 2017). Based on data from the Paris Biota and coeval beds in neighboring sites in the Bear Lake area (Fig. 1), we document the first reported occurrence of the ammonoids Tirolites harti, Tirolites aff. cassianus and Bajarunia cf. pilata, as well as a new genus Caribouceras nov. gen. and two new species Caribouceras slugense nov. gen., nov. sp. and Albanites americanus nov. sp. from the earliest Spathian platy shale and siltstone interval underlying the Middle Limestone unit and coeval beds. The Middle Limestone unit and coeval beds, which contains Tirolites harti, has been commonly considered as marking the beginning of the early Spathian in southeastern Idaho. This unit is overlain by the classic early Spathian assemblage of Columbites parisianus and Tirolites smithi, both species being common in the Bear Lake area. These new ammonoid occurrences in the Lower Shale unit or coeval beds, therefore, necessitate a downward revision of the Smithian/Spathian boundary (SSB) in the region and also modify the assumed correlation among sections in this area.

Section snippets

Ammonoids

After the severe late Smithian extinction (Tozer, 1982, Brayard et al., 2006, Jattiot et al., 2016), the appearance of the ammonoid genus Tirolites is consensually acknowledged as characterizing the early Spathian worldwide (Balini et al., 2010, Jenks et al., 2015, Shigeta and Nakajima, 2017). This certainly was the case for the western USA basin (e.g., Silberling and Tozer, 1968), based on data from ammonoids collected in the Bear Lake area by Hyatt and Smith (1905) and Smith (1932), and then

Exposures and sampled material

Early Triassic marine exposures of the western USA basin mainly consist of shale–limestone alternations of the Thaynes Group (sensu Lucas et al., 2007). These sediments were deposited over a large area covering western Wyoming, southern Montana, Idaho, Utah, and Nevada. They represent epicontinental marine environments of the Sonoma Foreland Basin, which was located within the northern intertropical zone on the western margin of Pangea during the Permian–Triassic transition (Dickinson, 2013).

Systematic paleontology

Ammonoid systematic descriptions follow the classification of Tozer (1981), and that further refined by Guex et al., 2005a, Guex et al., 2005b, Guex et al., 2010 and Jenks et al. (2013). Nautiloid systematic descriptions mainly follow the classifications of Sobolev (1989) and Shigeta et al. (2014). Exact quantitative morphological ranges for all ammonoid taxa are not permitted because of the compressed preservation of many specimens. We thus mainly report estimates for the diameter (D), whorl

Discussion

Extensive sampling of the studied outcrops clearly indicates that the lower Spathian deposits of the Lower Shale unit in Bear Lake County and coeval rocks in Caribou County (i.e., below the Tirolites [sensu Smith, 1932] and Columbites beds; Fig. 2) contain Tirolites harti, Bajarunia cf. pilata, and Albanites americanus nov. sp. The co-occurrence of T. harti and B. cf. pilata is common. At Dry Ridge, only B. cf. pilata has been found in the very base of the Spathian. At Schmid Ridge, two other

Acknowledgements

This work is a contribution to the ANR project AFTER (ANR-13-JS06-0001-01) and was also supported by the French “Investissements d’Avenir” program, project ISITE-BFC (ANR-15-IDEX-03). We acknowledge the Bear Lake County road department and private land owners D. M. Clow (Ogden, Utah), L. Rasmussen (Grace, Idaho) and his gate-keeper L. Hildreth (Soda Springs, Idaho) for allowing access to studied exposures. We are grateful to M. Balini and Y. Zakharov for helpful discussions. S. Lucas and M.

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      This therefore probably suggests a rather rapid restoration of regional communities after the LSC. Ammonoids and conodonts also show a rather rapid diversification during the early Spathian (Orchard, 2007; Brayard et al., 2009; Brayard et al., 2019). In addition, the Paris Biota from the earliest Spathian of southeastern Idaho highlights the existence of highly diversified and complex marine ecosystems in the immediate aftermath of the LSC (Brayard et al., 2017).

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    Corresponding editor: Gilles Escarguel.

    1

    The Paris Biota Team includes, in alphabetical order: Arnaud Brayard (team leader; Biogéosciences, Univ. Bourgogne Franche-Comté, Dijon, France), Kevin G. Bylund (Spanish Fork, UT, USA), Gilles Escarguel (LEHNA, Univ. Lyon 1, Lyon, France), Emmanuel Fara (Biogéosciences, Univ. Bourgogne Franche-Comté, Dijon, France), Nicolas Goudemand (IGFL, ENS-Lyon, Lyon, France), Pierre Gueriau (Synchrotron SOLEIL, Gif-sur-Yvette, France/Institute of Earth Sciences, University of Lausanne), James F. Jenks (West Jordan, UT, USA), L.J. Krumenacker (Idaho Museum of Natural History, Idaho State Univ., Pocatello, ID, USA), Nicolas Olivier (Magmas et Volcans, Univ. Clermont Auvergne, Clermont-Ferrand, France), Daniel A. Stephen (Dpt. of Earth Sciences, Utah Valley Univ., Orem, UT, USA), Christophe Thomazo (Biogéosciences, Univ. Bourgogne Franche-Comté, Dijon, France), Mathieu Thoury (IPANEMA, Univ. Paris-Saclay/Synchrotron SOLEIL, Gif-sur-Yvette, France), and Emmanuelle Vennin (Biogéosciences, Univ. Bourgogne Franche-Comté, Dijon, France).

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