Abnormal segregation for waxy and apiculus coloration, which is explained by assuming the presence of a gametophyte gene, ga linked with wx and C, was described. The irregularity of waxy segregation in these studies was first observed in the F₂ of a cross between a non-waxy progeny (wx+wx+1g+1g+C+C+) from plants exposed to the atomic bomb in Nagasaki, and a linkage tester characterized by waxy, Iiguleless and apiculus coloration (wx wxlg lg C C) (Table 1).