Skip to main content Skip to main navigation menu Skip to site footer
Type: Article
Published: 2019-11-26
Page range: 501–534
Abstract views: 85
PDF downloaded: 10

Taxonomic and nomenclatorial revision within the Neotropical genera of the subtribe Odontocheilina W. Horn in a new sense—22. Genus Cheilonycha Lacordaire, 1842 (Coleoptera: Cicindelidae)

Mendel University in Brno, Department of Forest Ecology, Zemědělská 3, 613 00 Brno, Czech Republic.
Coleoptera Cicindelidae Odontocheilina taxonomy Cheilonycha new species new subspecies distribution termitophilous habitat Neotropical Region

Abstract

Taxonomic and nomenclatorial revision of the genus Cheilonycha Lacordaire, 1842 with type species Cheilonycha chalybea (by original monotypy), based on Cicindela chalybea Dejean, 1825, is presented. It is concluded here that a subsequent type designation by Horn (1910), who without any explanation mentioned Ch. auripennis (Lucas, 1857) as the type species of the genus, was unjustified and must be considered invalid according to ICZN (1999). Examinations of type specimens have revealed that Cheilonycha auripennis sensu auctorum (sensu Horn 1922 and subsequent authors) is a complex of taxa. As a result, one new subspecies of Ch. auripennis (Lucas, 1857) and one new species of the genus are described and the following taxa of Cheilonycha are recognized and presented here: Ch. chalybea (Dejean, 1825), Ch. a. auripennis (Lucas, 1857), Ch. a. chiquitosiana ssp. nov. and Ch. bucephalauripennis sp. nov. Type specimens of Ch. a. angustedilatata (W. Horn, 1922) based on Odontochila (Chilonycha) auripennis angustedilatata W. Horn, 1922, proved to be fully conspecific with the type specimens of Ch. auripennis based on Cicindela auripennis Lucas, 1857. Consequently, Ch. a. angustedilatata syn. nov. is treated here as a junior synonym of Ch. a. auripennis. The history of the taxonomy of these taxa, their distribution and biology, including photographs of the termitophilous habitat, relevant lectotype designations, descriptions (redescriptions respectively) and illustrations in colour photographs of the habitus, diagnostic characters and variability are provided.

 

References

  1. Agassiz, L. (1846) Nomenclatoris Zoologici, Index Universalis, continens nomina systematica classium, ordinum, familiarum et generum animalium mnium, tam viventium quam fossilium, secundum ordinem alphabeticum unicum disposita, adjectis homonymiis plantarum, nec non variis adnotationibus et emendationibus. Jent et Gassmann, Soloduri, viii + 393 pp.

    Arndt, E., Paarmann, W. & Adis, J. (1996) Description and key of larval Cicindelidae from Brazil (Coleoptera: Caraboidea). Acta Societatis Zoologicae Bohemicae, 60, 293–315.

    Bates, H.W. (1872) Notes on Cicindelidae and Carabidae, and descriptions of new species (No. 14). Entomologist’s Monthly Maazine, 8, 263–266.

    Berg, K. (1900) Termitariophile. Commucaciones del Museo nacionalde Buenos Aires, 1, 212–2015.

    Blackwelder, R.E. (1944) Checklist of the coleopterous insects of Mexico, Central America, The west Indies, and South America Part 1. Smithsonian Institution United States National Museum, Bulletin, 185, 1–188.

    https://doi.org/10.5479/si.03629236.185.i

    Bousquet, Y. (2002) Additions and corrections to the world catalogue of genus-group names of Geadephaga (Coleoptera) published by Wolfgang Lorenz (1998). Folia Heyrovskyana, Supplementum 9, 1–78.

    Bruch, C. (1911) Catálogo sistemático de los coleópteros de la República Argentina, pars 1, Familia Carabidae (Cicindelinae, Carabinae). Revista del Museo de La Plata, 17 (4), 143–180.

    https://doi.org/10.5962/bhl.title.8799

    Cassola, F. (2001) Studies of tiger beetles. CXXIII. Preliminary approach to the macrosystematics of the tiger beetles (Coleoptera, Cicindelidae). Russian Entomology Journal, 10 (3), 265–272.

    Cassola, F. & Pearson, D.L. (2001) Neotropical Tiger Beetles (Coleoptera: Cicindelidae): Checklist and Biogeography. Biota Colombiana, 2 (1), 3–24.

    Dejean, P.M.F.A. (1825) Species général des Coléoptères de la collection de M. le Comte Dejean. Tome premier. Crevot, Paris, xxx + 463 pp.

    https://doi.org/10.5962/bhl.title.2086

    Dokhtouroff, W. (1887) Matériaux pour servir a l’étude des Cicindèlides. Annales de la Société Entomologique de Belgique, 31, 152–158.

    Döbler, H. (1973) Katalog der in den Sammlungen des ehemaligen Deutschen Entomologischen Institutes aufbewahrten Typen—IX. Coleoptera, Cicindelidae. Beiträge zur Entomologie, 23, 355–419.

    Duran, D.P. & Moravec, J. (2013) A new species of the genus Pentacomia from Panama (Coleoptera: Cicindelidae) Acta Entomologica Musei Nationalis Pragae, 53, 49–57.

    Erwin, T.L. & Pearson, D.L. (2008) A treatise on the Western Hemisphere Caraboidea (Coleoptera). Their classification, distributions, and ways of the life. Vol. II. Carabidae—Nebriformes 2—Cicindelitae. Pensoft Series Faunistica 84. Pensoft Publishers, Sofia, 366 pp., 33 pls.

    Fleutiaux, E. (1892) Catalogue systematique des Cicindelidae. Imprimerie H. Vaillant-Carmanne, Liege, 186 pp.

    https://doi.org/10.5962/bhl.title.48634

    Guerra, J.F. (1993) Some observations of the Termite mound-dvelling tiger beetle Cheilonycha auripennis Lucas , from northeastern Bolivia. Cicindela, 25, 23–26.

    Harris, R.A. (1979) A glossary of surface sculpturing. In: Andrews, G. (Ed.), Department of Food and Agriculture Division of Plant Industry, Sacramento. Occasional papers of Laboratory Services / Entomology, 28, 1–31.

    Horn, W. (1899) Ueber das System der Cicindeliden. Deutsche Entomologische Zeitschrift, 1, 33–51.

    Horn, W. (1910) Coleoptera Adephaga, Fam. Carabidae, Subfam. Cicindelinae. In: Wytsman, P. Genera Insectorum, 82B, 103–208, pls. 6–15.

    Horn, W. (1915) Coleoptera Adephaga, Fam. Carabidae, Subfam. Cicindelinae. In: Wytsman, P., Genera Insectorum, 82C, 209–487, pls. 16–23.

    Horn, W. (1922) Studien über neue und alte Cicindeliden (Col.), (Neubeschreibungen, Synonymie, Faunistik). Zoologische Mededeelingen, 7, 90–112.

    Horn, W. (1924) Les Cicindelides des “Vaccarias”du Rio Vaccaria, dans le Matto Grosso (Brésil) et leur termitophilie. Bulletin de la Société de Cluj, 2, 46–48.

    Horn, W. (1926a) Carabidae, Cicindelinae. In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus. Junk, Berlin, 345 pp.

    Horn, W. (1926b) Ueber die Genese von Cicindela hybrida- und campestris-Formen, sowie die Cicindeliden-Fauna von Uganda und den “Vacarias” von Matto Grosso. Entomologische Mitteilungen, 15 (1), 69–78.

    Horn, W. (1931) Die Cicindelidenfauna von Bolivien. Revista de Entomologia, 1 (1), 19–30.

    Horn, W. (1934) On the three forms of Odontochila subgenus Chilonycha (Cicind., Col). Extracto de la Revista Chilena de Historia Natural, 38, 124–125.

    Horn, W. (1938) 2000 Zeichnungen von Cicindelinae. Entomologische Beihefte aus Berlin-Dahlem, 5, 1–71, 90 Tafeln.

    ICZN (1999) International Code of Zoological Nomenclature, fourth edition, adopted by the International Union of Biological Sciences. International Trust for Zoological Nomenclature, London, xxix + 306 pp.

    Kippenhan, M.G. (1997) A review of the Neotropical tiger beetle genus Oxygonia Mannerheim (Coleoptera: Cicindelidae). Contributions on Entomology, International 2 (3), 303–353.

    Lacordaire, J.T. (1842) Révision de la famille des Cicindélides de l’ordre des Coléoptères. Quadart, Liége, 40 pp. [published in 1843 in Mémoires de la Société Royale des Sciences de Liége, 1, 85–118, however, reprints Quadart, Liége appeared in 1842]

    Lacordaire, J.T. (1843) Révision de la famille des Cicindélides (Cicindelidae) de l’ordre des Coléoptères, accompagnée de la création de quelques genres nouveaux Mémoires de la Société Royale des Sciences de Liége, 1, 85–118.

    Laporte de Castelnau, F.L. (1834) Observations sur la tribu des Cicindélètes. Revue Entomologique, 2, 27–38.

    Latreille, P.A. (1802) Histoire naturelle générale et particulière des Crustacés et des Insectes, Paris, 3, 79. Ouvrage faisant suite à l’ histoire naturelle générale et particulière, compose par Leclere de Buffon, et rédigée par C. S. Sonnini, member de plusieurs sociètes savants. Familles naturelles des genres. Tome Troisième. Dufart, Paris, 448 pp. [pp. i–xii + pp. 13–447 + (1)]

    https://doi.org/10.5962/bhl.title.15764

    Linard, B., Cramton-Platt, A., Moriniere, J., Timmersmans, M.J.T.N., Andújar, C., Arribas, P., Miller, K.E., Lipecki, J., Favreau, E., Hunter, A., Goméz-Rodríguez, C., Barton, C., Nie, R., Gillett, C.P.D.T., Breeschoten, T., Bocák, L. & Vogler, A.P. (2018) The contribution of mitochondrial metagenomics to large-scale data mining and phylogenetic analysis of Coleoptera. Molecular Phylogenetics and Evolution, 128 (2018), 1–11.

    https://doi.org/10.1016/j.ympev.2018.07.008

    Lopéz-Lopéz, A. & Vogler, A. (2017) The mitogenome phylogeny of Adephaga (Coleoptera). Molecular Phylogenetics and Evolution, 7 (114), 166–174. https://doi.org/10.1016/j.ympev.2017.06.009

    Lorenz, W. (1998a) Systematic list of extant ground beetles of the world (Insecta, Coleoptera, “Geadephaga”: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Privately published, Tutzing, 502 pp.

    Lorenz, W. (1998b) Nomina Carabidarum: a directory of the scientific names of ground beetles (Insecta,Coleoptera, “Geadephaga” : Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Privately published, Tutzing, 937 pp.

    Lorenz, W. (2005a) Systematic list of extant ground beetles of the world (Insecta, Coleoptera, “Geadephaga”:Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second edition. Wolfgang Lorenz, Tutzing, iv + 530 pp.

    Lorenz, W. (2005b) Nomina Carabidarum: a directory of the scientific names of ground beetles (Insecta,Coleoptera, “Geadephaga”: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). 2nd Edition. Wolfgang Lorenz, Tutzing, 937 pp.

    Lucas, P.H. (1857–1859) Animaux nouveaux ou rares recueillis pendant l’expédition dans les parties centrales de l’Amérique du Sud, de Rio de Janeiro a Lima, et de Lima au Para. Entomologie. P. Bertrand, Paris, 204 pp., 20 pls.

    Mandl, K. (1958) Beiträge zur Kenntnis der Insektenfauna Boliviens 5. Coleoptera 4, Cicindelidae. Opuscula Zoologica, 11, 1–12.

    Mannerheim, M. (1837) Mémoire sur Quelques genres et espèces de Carabiques. Bulletin de la Société Impériale des Naturalistes de Moscow, 2, 1–49.

    Moravec, J. (2002) Tiger beetles of Madagascar 2. A monograph of the genus Physodeutera (Coleoptera: Cicindelidae). Nakladatelství Kabourek, Zlín, 290 pp.

    Moravec, J. (2007) Tiger beetles of Madagascar 1. A monograph of the genus Pogonostoma (Coleoptera: Cicindelidae). Nakladatelství Kabourek, Zlín, 499 pp.

    Moravec, J. (2010) Tiger beetles of the Madagascan Region (Madagascar, Seychelles, Comoros, Mascarenes, and other islands. Taxonomic revision of the 17 genera occurring in the region (Coleopterea: Cicindelidae). Biosférická rezervace Dolní Morava, o.p.s., Lednice na Moravě, 429 pp.

    Moravec, J. (2012a) Taxonomic and nomenclatorial revision within the Neotropical genera of the subtribe Odontochilina in a new sense—1. Some changes in taxonomy and nomenclature within the genus Odontocheila (Coleoptera: Cicindelidae). Acta Musei Moraviae, Scientiae biologicae, 97 (2), 13–33.

    Moravec, J. (2012b) Taxonomic and nomenclatorial revision within the Neotropical genera of the subtribe Odontochilina W. Horn in a new sense—2. Brzoskaicheila gen. nov., a new genus for Cicindela hispidula Bates, 1872, and Brzoskaicheila crassisculpta sp. nov. (Coleoptera: Cicindelidae). Acta Musei Moraviae, Scientiae biologicae, 97 (2), 35–48.

    Moravec, J. (2012c) Taxonomic and nomenclatorial revision within the Neotropical genera of the subtribe Odontochilina W. Horn in a new sense—3. Pentacomia (Mesacanthina) punctum (Klug) and P. (M.) ronhuberi sp.nov. (Coleoptera: Cicindelidae). Acta Musei Moraviae, Scientiae biologicae, 97 (2), 49–63.

    Moravec, J. (2013) Taxonomic and nomenclatorial revision within the Neotropical genera of a subtribe Odontochilina W. Horn in a new sense—4. A new species and a new synonymy within the genus Odontocheila. (Coleoptera: Cicindelidae). Acta Musei Moraviae, Scientiae biologicae, 98 (1), 53–73.

    Moravec, J. (2014) Taxonomic and nomenclatorial revision within the Neotropical genera of the subtribe Odontocheilina W. Horn in a new sense—9. Odontocheila pentacomioides W. Horn, 1900 comb. restit.; O. cyanella pseudomargineguttata W. Horn, 1930 syn. nov., a junior synonym of O. spinipennis Chaudoir, 1843. Acta Musei Moraviae, Scientiae biologicae, 99 (1), 47–64.

    Moravec, J. (2015a) Taxonomic and nomenclatorial revision within the Neotropical genera of the subtribe Odontochilina W. Horn in a new sense—11. The genus Cenothyla Rivalier, 1969 (Coleoptera: Cicindelidae). Studies and Reports Taxonomical Series, 11 (1), 77– 122.

    Moravec, J. (2015b) Taxonomic revision within the Neotropical genus Oxygonia Mannerheim—1 (Coleoptera: Cicindelidae). Folia Heyrowskyana, Series A, 23 (2), 27–70.

    Moravec, J. (2016a) Taxonomic and nomenclatorial revision within the Neotropical genera of the subtribe Odontochilina W. Horn in a new sense—15. The genus Opisthencentrus W. Horn (Coleoptera: Cicindelidae). Zootaxa, 4097 (3), 332–340. https://doi.org/10.11646/zootaxa.4097.3.2

    Moravec, J. (2016b) Taxonomic and nomenclatorial revision within the Neotropical genera of the subtribe Odontochilina W. Horn in a new sense—16. Pentacomia (Mesochila) procera (Chaudoir), P. (M.) conformis (Dejean), and P. (M.) proceroides sp. nov. (Coleoptera: Cicindelidae). Zootaxa 4127 (2), 276–300.

    https://doi.org/10.11646/zootaxa.4127.2.3

    Moravec, J. (2016c) Taxonomic and nomenclatorial revision within the Neotropical genera of the subtribe Odontocheilina W. Horn in a new sense—17. O. cajennensis species-complex and key to species of Odontocheila cajennensis species-group (Coleoptera: Cicindelidae). Acta Musei Moraviae, Scientiae biologicae, 101 (1), 7–53, 2016

    Moravec, J. (2017) Taxonomic revision of the Neotropical genus Oxygonia Mannerheim—2 Coleoptera: Cicindelidae). Folia Heyrovskyana, Series A, 25 (2), 31–122.

    Moravec, J. (2018a) Taxonomic and nomenclatorial revision within the Neotropical genera of the subtribe Odontocheilina W. Horn in a new sense—20. Beckerium W. Horn stat. restit., Mesochila Rivalier stat. nov. and Poecilochila Rivalier stat. nov. (Coleoptera: Cicindelidae). Acta Musei Moraviae, Scientiae biologicae, 103 (2), 127–206

    Moravec, J. (2018b) Taxonomic revision of the Neotropical tiger beetle genera of the subtribe Odontocheilina. Volume 1. Odontocheila Laporte de Castelnau, Cenothyla Rivalier and Phyllodroma Lacordaire. (Coleoptera: Cicindelidae). Biosférická rezervace Dolní Morava, o.p.s., Lednice na Moravě, 623 + 2 pp.

    Moravec, J. & Brzoska, D. (2013) Taxonomic and nomenclatorial revision within the Neotropical genera of the subtribe Odontochilina W. Horn in a new sense—5. A new species of the genus Pentacomia from Costa Rica. Acta Musei Moraviae, Scientiae biologicae, 98 (1), 75–84.

    Moravec, J. & Brzoska, D. (2014a) Taxonomic and nomenclatorial revision within the Neotropical genera of the subtribe Odontocheilina W. Horn in a new sense—7. Pentacomia (Pentacomia) davidpearsoni sp. nov., a new species from Bolivia related to P. (P.) speculifera (Brullé) (Coleoptera: Cicindelidae). Acta Musei Moraviae, Scientiae Biologicae, 9 (1), 15–33.

    Moravec, J. & Brzoska, D. (2014b) Taxonomic and nomenclatorial revision within the Neotropical genera of the subtribe Odontocheilina W. Horn in a new sense—8. Redescription and lectotype designation of Pentacomia (Pentacomia) lanei (W. Horn), with a new record from Paraguay. Acta Musei Moraviae, Scientiae Biologicae, 99 (1), 35–46.

    Moravec, J. & Brzoska, D. (2014c) Taxonomic and nomenclatorial revision within the Neotropical genera of the subtribe Odontocheilina W. Horn in a new sense—10. Odontocheila castelnaui species-group (Coleoptera: Cicindelidae). Studies and Reports Taxonomical Series, 10 (2), 495–526.

    Moravec, J. & Brzoska, D. (2015) Taxonomic and nomenclatorial revision within the Neotropical genera of the subtribe Odontocheilina W. Horn in a new sense—12. Odontocheila angelsolisi spec. nov., O. mirekskrabali sp. nov. and related species of a newly proposed Odontocheila cajennensis species-group (Coleoptera: Cicindelidae). Acta Musei Moraviae, Scientiae biologicae, 100 (1), 23–66.

    Moravec, J. & Brzoska, D. (2019) Taxonomic and nomenclatorial revision within the Neotropical genera of the subtribe Odontocheilina W. Horn in a new sense—21. Pentacomia paranigrimarginata sp. nov. and P. nigrimarginata Huber with revised key to Pentacomia species (Coleoptera: Cicindelidae). Zootaxa, 4612 (1), 71–84.

    https://doi.org/10.11646/zootaxa.4612.1.4

    Moravec, J. & Duran, D.P. (2013) Taxonomic and nomenclatorial revision within the Neotropical genera of the subtribe Odontochilina W. Horn in a new sense. 6. Odontocheila fraternum sp. nov., a new species sister to O. gilli (Coleoptera: Cicindelidae). Acta Entomologica Musei Nationalis Pragae, 53, 585–599.

    Moravec, J. & Huber, R. (2015) Taxonomic and nomenclatorial revision within the Neotropical genera of the subtribe Odontocheilina W. Horn in a new sense. 13. The genus Mesacanthina Rivalier, stat. nov., separated from the genus Pentacomia Bates (Coleoptera: Cicindelidae). Acta Musei Moraviae, Scientiae biologicae, 100 (1), 67–114.

    Moravec, J., Brzoska, D. & Huber, R.L. (2017) Taxonomic and nomenclatorial revision within the Neotropical genera of the subtribe Odontocheilina W. Horn in a new sense—18. Six Mexican and Central American species related to Odontocheila mexicana Laporte de Castelnau and O. ignita Chaudoir, with a description of O. potosiana sp. nov. (Coleoptera: Cicindelidae). Zootaxa, 4231 (4), 451–499.

    https://doi.org/10.11646/zootaxa.4231.4.1

    Moravec, J., Huber, R. & Brzoska, D. (2017) Taxonomic and nomenclatorial revision within the Neotropical genera of the subtribe Odontocheilina W. Horn in a new sense—19. Odontocheila microptera nom. nov., a new replacement name for O. euryoides W. Horn, 1922, and lectotype designation of O. nitidicollis (Dejean, 1825) (Coleoptera: Cicindelidae). Zootaxa, 4347 (1), 109–127.

    https://doi.org/10.11646/zootaxa.4347.1.6

    Moravec, J., Huber, R.L. & Dheurle, C. (2015) Taxonomic and nomenclatorial revision within the Neotropical genera of the subtribe Odontocheilina W. Horn in a new sense—14. Pentacomia (Pentacomia) chrysammoides sp. nov., a new species related to P. chrysamma (Coleoptera: Cicindelidae). Acta Musei Moraviae, Scientiae biologicae, 100 (2), 217–249.

    Moravec, J. & Kudrna, A. Jnr. (2002) Ronhuberia gen n. with type species Pentacomia fernandezi (Cassola) comb. n.; R. eurytarsipennis (W. Horn) comb. n. Cicindela, 34 (3–4), 17–37.

    Naviaux, R. (2002) Faune du Brésil. Description de deux nouveaux Ctenostoma, catalogue des Cicindèles (Coleoptera, Cicindelidae). Bulletin de la Société entomologique de France, 107 (3), 265–273.

    Nichols, S.W. (1989) The Torre-Bueno glossary of entomology, revised edition of Torre Bueno (Rollin, J.) (1937) A glossary of entomology including Tulloch G. S. 1962: Supplement A. The New York Entomological Society, American Museum of Natural History, New York, 840 pp.

    Pearson, D.L. & Vogler, A.P. (2001) Tiger beetles, the evolution, ecology and diversity of the Cicindelids. Cornell University Press, Ithaca and London, 333 pp.

    Pearson, D.L. Guerra, J.F. & Brzoska, D.W. (1999) The Tiger beetles of Bolivia: their Identification, Distribution and Natural History (Coleoptera: Cicindelidae). Contributions on Entomology, International, 3 (4), 379–524.

    Pražák, J.M., Novotný, M. & Sedláček, J. (1955) Latinsko-Český Slovník. Part A–Z. Sátní pedagogické nakladatelství, Praha, 743 pp. & 684 pp. [Part A–K: 743 pp., Part L–Z: 684 pp.]

    Putchkov, A.V. & Cassola, F (2005) Tiger beetles deserve separate family status in suborder Adephaga (Coleoptera, Cicindelidae). Bulletin de la Société entomologique de France, 110, 281–293.

    Reiche, M.L. (1842) Colèoptères de Colombie. Revue Zoologique, par la Société cuvierienne, 5, 238–242.

    Reichardt, H. (1977) A synopsis of the genera of Neotropical Carabidae (Insecta: Coleoptera). Questiones Entomologicae, 13, 346–493.

    Rivalier, E. (1969) Démembrement du genre Odontochila (Col. Cicindelidae) et revision des principales espèces. Annales de la Société entomologique de France, Nouvelle Série, 5 (1), 195–237.

    Rivalier, E. (1971) Remarques sur la tribu des Cicindelini (Col. Cicindelidae) et sa subdivision en sous-tribus. Nouvelle Revue d’Entomologie, 1, 135–143.

    Roza, A.S. & Mermudes, J.R.M. (2015) Description of the immature stages of Pentacomia (Mesochila) smaragdula (Dejean 1825) (Coleoptera: Carabidae, Cicindelinae), with notes on the species distribution. Zootaxa, 4057 (3), 399–408.

    http://dx.doi.org/10.11646/zootaxa.4057.3.6

    Roza, A.S. & Mermudes, J.R.M. (2017) Distribution of Pentacomia (Mesochila) Rivalier, 1969 (Coleoptera: Carabidae, Cicindelinae ). Transactions American Entomological Society, TAES, 143, 601–623.

    https://doi.org/10.3157/061.143.0305

    Sawada, H. & Wiesner, J. (2002) The Tiger Beetles of Paraguay (Insecta: Coleoptera: Cicindelidae), 67th. Contribution towards the knowledge of Cicindelidae. Coleoptera, Schwanfelder Coleopterologische Mitteilungen, 4 (2000), 285–325.

    Schade, F. (1933) Beitrag zur Kunde der Cicindeliden Zentral-Paraguays. Entomologische Rundschau, 50, 248–251.

    Vogler, A.P. & Pearson, D.L. (1996) A molecular phylogeny of the tiger beetles (Cicindelidae): Congruence of mitochondrial and nuclear rDNA datasets. Molecular Phylogenetics and Evolution, 6, 321–338.

    https://doi.org/10.1006/mpev.1996.0083

    Wasmann, E. (1895a) Ueber termitophile Cicindeliden. Deutsche Entomologische Zeitschrift, 2, 289–290.

    Wasmann, E. (1895b) Die Ameisen– und Termitengäste von Brasilien. Verhandlungen des Zoologisch-Botanischen Vereins in Wien, 45, 137–145.

    Wiesner, J. (1992) Verzeichnis der Sandlaufkäfer der Welt. Checklist of the tiger beetles of the world (Coleoptera, Cicindelidae). Verlag Erna Bauer, Keltern, 364 pp.

    Wiesner, J. & Bandinelli, A. (2014) Notes on the tiger beetles (Coleoptera: Cicindelidae) of Argentina. 117. Contribution towards the knowledge of Cicindelidae. Insecta Mundi, 0377, 1–66.