* Corresponding authors

^a Department of Biochemistry, University of Wisconsin-Madison, 440 Henry Mall, Madison, WI 53706, USA
E-mail: weibel@biochem.wisc.edu

^b Department of Chemical and Biological Engineering, University of Wisconsin-Madison, 1415 Engineering Dr., Madison, WI 53706, USA

^c Department of Mathematics, University of Wisconsin-Madison, 480 Lincoln Dr., Madison, WI 53706, USA
E-mail: spagnolie@math.wisc.edu

^d Department of Chemistry, University of Wisconsin-Madison, 1101 University Ave., Madison, WI 53706, USA

^e Department of Biomedical Engineering, University of Wisconsin-Madison, 1550 Engineering Dr., Madison, WI 53706, USA