Hostname: page-component-8448b6f56d-c4f8m Total loading time: 0 Render date: 2024-04-19T21:24:08.583Z Has data issue: false hasContentIssue false

Selected Ordovician Trilobites from the Lake St. John District of Quebec and Their Bearing on Systematics

Published online by Cambridge University Press:  11 August 2017

Pierre J. Lespérance
Affiliation:
Département de géologie, Université de Montréal, P.O. Box 6128, Montreal, Canada, H3C 3J7
Sylvain Desbiens
Affiliation:
Geological Survey of Canada, 601 Booth Street, Ottawa, Canada, K1A 0E8

Abstract

The thorax of Hypodicranotus has ten segments and a spine on the eighth. The ages of Erratencrinurus s.l. spicatus and Erratencrinurus (Erratencrinurus?) vigilans in the Lake St. John district do not confirm their temporal roles leading to subgenera of Erratencrinurus, as has been recently suggested. Phylogenetic analyses of large data sets of species previously referred to Encrinuroides and Physemataspis yield a minimal length cladogram containing 18 species. Encrinuroides is restricted to four species, two of which have biogeographic affinities with Iapetus. These results lead to three clades, named the Walencrinuroides n. gen. clade, Frencrinuroides n. gen. clade, and finally the Physemataspis clade, with an enlarged concept of the genus with the erection of Physemataspis (Prophysemataspis) n. subgen. These last three clades are restricted to North America and Scotland, with alternating predominance of one region. Walencrinuroides s.l. gelaisi n. gen. n. sp. is described. New morphological data on Erratencrinurus s.l. spicatus confirm its close relationship with the clades discussed above. Data are insufficient for phylogenetic analysis of selected cheirurine species here surveyed. Eye position, glabellar segmentation, and pygidial shape differentiate the genera Ceraurus and Gabriceraurus; emended diagnoses of these genera are presented. Ceraurus globulobatus and C. matranseris are distinct, but morphologically close to one another. The status of Gabriceraurus dentatus can be stabilized on its extant types.

Type
Research Article
Copyright
Copyright © 1995 by The Paleontological Society, Inc. 

Access options

Get access to the full version of this content by using one of the access options below. (Log in options will check for institutional or personal access. Content may require purchase if you do not have access.)

References

Angelin, N. P. 1854. Palaeontologia Scandinavica. Pars I, Iconographia Crustaceorum formationis transitionis, Fasciculum II, p. iix, 21–92, PL 25–41, Lund.Google Scholar
Barnes, C. R., Norford, B. S., and Skevington, D. 1981. The Ordovician System in Canada. Correlation chart and explanatory notes. International Union of Geological Sciences, Publication 8, 27 p.Google Scholar
Barton, D. C. 1913. A new genus of the Cheiruridae, with description of some new species. Bulletin of the Museum of Comparative Zoology at Harvard College, 54:545556.Google Scholar
Bergström, S. M., and Mitchell, C. E. 1986. The graptolite correlation of the North American Upper Ordovician standard. Lethaia, 19:247266.CrossRefGoogle Scholar
Bolton, T. E. 1966. Catalogue of type invertebrate fossils of the Geological Survey of Canada. Volume 3. Geological Survey of Canada, 203 p.CrossRefGoogle Scholar
Bradley, J. H. Jr. 1930. Fauna of the Kimmswick Limestone of Missouri and Illinois. Contributions from Walker Museum, 2:i–v, 219290.Google Scholar
Burskyi, N. N. 1966. Encrinuridae from the Ordovician deposits of North Pai Khoi and Vaigach Island. Nauchno-issledovatel'skij Institut Geologii Arktiki, Uchenye Zapiski, 11:7984, Leningrad [in Russian].Google Scholar
Caster, K. E., Dalvé, E. A., and Pope, J. K. 1961. Elementary guide to the fossils and strata of the Ordovician in the vicinity of Cincinnati, Ohio. Cincinnati Museum of Natural History, 47 p.Google Scholar
Chatterton, B. D. E., and Ludvigsen, R. 1976. Silicified Middle Ordovician trilobites from the South Nahanni River area, District of Mackenzie, Canada. Palaeontographica, Abteilung A, 154:1106.Google Scholar
Cooper, B. N. 1953. Trilobites from the Lower Champlainian Formations of the Appalachian Valley. Geological Society of America Memoir, 55, 69 p.Google Scholar
Cooper, G. A., and Kindle, C. H. 1936. New brachiopods and trilobites from the Upper Ordovician of Perce, Quebec. Journal of Paleontology, 10:348372.Google Scholar
Dean, W. T. 1973. Lower Ordovician trilobites from the Summerford Group at Virgin Arm, New World Island, northeastern Newfoundland. Geological Survey of Canada Bulletin, 240, 43 p.Google Scholar
Dean, W. T. 1979. Trilobites from the Long Point Group (Ordovician), Port au Port Peninsula, southwestern Newfoundland. Geological Survey of Canada Bulletin, 290, 53 p.Google Scholar
DeMott, L. L., Edited by Sloan, R. E., Shaw, F. C., and Tripp, R. P. 1987. Platteville and Decorah trilobites from Illinois and Wisconsin, p. 6398. In Sloan, R. E. (ed.), Middle and Late Ordovician lithostratigraphy and biostratigraphy of the Upper Mississippi Valley. Minnesota Geological Survey, Report of Investigations 35.Google Scholar
Desbiens, S., and Lespérance, P. J. 1989. Stratigraphy of the Ordovician of the Lac Saint-Jean and Chicoutimi outliers, Quebec. Canadian Journal of Earth Sciences, 26:11851202.CrossRefGoogle Scholar
Edgecombe, G. D., and Chatterton, B. D. E. 1987. Heterochrony in the Silurian radiation of encrinurine trilobites. Lethaia, 20:337351.CrossRefGoogle Scholar
Edgecombe, G. D., and Chatterton, B. D. E., 1990. Systematics of Encrinuroides and Curriella (Trilobita), with a new Early Silurian encrinurine from the Mackenzie Mountains. Canadian Journal of Earth Sciences, 27:820833.CrossRefGoogle Scholar
Edgecombe, G. D., Speyer, S. E., and Chatterton, B. D. E. 1988. Protaspid larvae and phylogenetics of encrinurid trilobites. Journal of Paleontology, 62:779799.Google Scholar
Esker, G. E. III. 1964. New species of trilobites from the Bromide Formation (Pooleville Member) of Oklahoma. Oklahoma Geology Notes, 24:195209.Google Scholar
Evitt, W. R. 1953. Observations on the trilobite Ceraurus . Journal of Paleontology, 27:3348.Google Scholar
Evitt, W. R., and Tripp, R. P. 1977. Silicified Middle Ordovician trilobites from the families Encrinuridae and Staurocephalidae. Palaeontographica, Abteilung A, 157:109174.Google Scholar
Foerste, A. F. 1920. The Kimmswick and Plattin Limestones of Northeastern Missouri. Journal of the Scientific Laboratories of Denison University, 19:175224.Google Scholar
Forey, P. L., Humphries, C. J., Kitching, I. L., Scotland, R. W., Siebert, D. J., and Williams, D. M. 1992. Cladistics—a practical course in systematics. Systematics Association Publication 10, Clarendon Press, Oxford, 191 p.Google Scholar
Fortey, R. A. 1980. The Ordovician trilobites of Spitsbergen. III. Remaining trilobites of the Valhallfonna formation. Norsk Polarinstitutt, Skrifter Nr. 171, 163 p.Google Scholar
Frederickson, E. A. 1964. Two Ordovician trilobites from southern Oklahoma. Oklahoma Geology Notes, 24:7175.Google Scholar
Green, J. 1832. Monograph of the trilobites of North America. J. Brano, Philadelphia, 93 p.Google Scholar
Hall, J. 1847. Palaeontology of New-York. Volume I. Containing descriptions of the organic remains of the Lower Division of the New-York System. Albany, 338 p.Google Scholar
Hawle, I., and Corda, A. J. 1847. Prodrom einer Monographic der böhmischen Trilobiten. Abhandlungen Böhmischen Gesellschaft Wissenschaften Prague, 5:117292.Google Scholar
Hessin, W. A. 1988a. Leviceraurus, a new cheirurine trilobite from the Cobourg Formation (Middle–Upper Ordovician), Southern Ontario, Canada. Journal of Paleontology, 62:8793.CrossRefGoogle Scholar
Hessin, W. A. 1988b. Partial regeneration of a genal spine by the trilobite Ceraurus plattinensis . Lethaia, 21:285288.CrossRefGoogle Scholar
Hessin, W. A. 1989. Ceraurus and related trilobites from the Middle Ordovician Bobcaygeon Formation of south-central Ontario, Canada. Canadian Journal of Earth Sciences, 26:12031219.CrossRefGoogle Scholar
Hessler, R. R. 1962. Secondary segmentation in the thorax of trilobites. Journal of Paleontology, 36:13051312.Google Scholar
Johnson, T. T. 1985. Trilobites of the Thomas T. Johnson Collection—How to Find, Prepare and Photograph Trilobites. Dayton, Ohio, 178 p.Google Scholar
Krueger, H.-H. 1971. Encrinuriden aus ordovizischen Geschieben (Teil I). Geologie, 20:11321169.Google Scholar
Krueger, H.-H. 1972. Nachweis der trilobitengattung Raymondella in Geschieben. Geologie, 21:856858.Google Scholar
Lane, P. D. 1971. British Cheiruridae (Trilobita). Palaeontographical Society Monographs, 95 p.CrossRefGoogle Scholar
Lespérance, P. J., Malo, M., Sheehan, P. M., and Skidmore, W. B. 1987. A stratigraphical and faunal revision of the Ordovician–Silurian strata of the Perce area, Quebec. Canadian Journal of Earth Sciences, 24:117134.CrossRefGoogle Scholar
Lespérance, P. J., and Tripp, R. P. 1985. Encrinurids (Trilobita) from the Matapédia Group (Ordovician), Perce, Quebec. Canadian Journal of Earth Sciences, 22:205213.CrossRefGoogle Scholar
Logan, W. E. sir, Murray, A., Hunt, T. S., and Billings, E. 1863. Report of Progress from its commencement to 1863 (Geology of Canada). Geological Survey of Canada, 983 p.CrossRefGoogle Scholar
Ludvigsen, R. 1976. New cheirurinid trilobites from the lower Whittaker Formation (Ordovician), southern Mackenzie Mountains. Canadian Journal of Earth Sciences, 13:947959.CrossRefGoogle Scholar
Ludvigsen, R. 1977. The Ordovician trilobite Ceraurinus Barton in North America. Journal of Paleontology, 51:959972.Google Scholar
Ludvigsen, R. 1978. Towards an Ordovician trilobite biostratigraphy of Southern Ontario. Michigan Basin Geological Society, Special Paper, 3:7384.Google Scholar
Ludvigsen, R. 1979a. Fossils of Ontario. Part 1: The Trilobites. Royal Ontario Museum, Life Sciences Miscellaneous Publications, 96 p.Google Scholar
Ludvigsen, R. 1979b. A trilobite zonation of Middle Ordovician rocks, Southwestern District of Mackenzie. Geological Survey of Canada Bulletin, 312, 99 p.Google Scholar
Ludvigsen, R., and Chatterton, B. D. E. 1991. The peculiar Ordovician trilobite Hypodicranotus from the Whittaker Formation, District of Mackenzie. Canadian Journal of Earth Sciences, 28:616622.CrossRefGoogle Scholar
Miller, S. A., and Gurley, W. F. E. 1894. Description of some new species of invertebrates from the Palaeozoic rocks of Illinois and adjacent states. Illinois State Museum of Natural History Bulletin, 3, 81 p.Google Scholar
Owen, A. W., and Heath, R. A. 1990 [1989]. A revision of the upper Ordovician trilobite genus Erratencrinurus with a description of a new species from Hadeland. Norsk Geologisk Tidsskrift, 69:225233.Google Scholar
Přibyl, A., and Vaněk, J. 1972. Hypodricranotinae nov. subfam. (Trilobita) aus dem mittleren Ordovicium von Nordamerika. Casopis pro mineralogii a geologii, 17:429430.Google Scholar
Přibyl, A., and Vaněk, J. 1985. Description of new cheirirud (sic) genera and subgenera, p. 157171. In Přibyl, A., Vaněk, J., and Pek, I., Phylogeny and taxonomy of Family Cheiruridae (Trilobita). Acta Universitatis Palackianae Olomucensis, Facultas Rerum Naturalium, Geographica–Geologica XXIV, 83:107–193.Google Scholar
Price, D. 1974. Trilobites from the Sholeshook Limestone (Ashgill) of South Wales. Palaeontology, 17:841868.Google Scholar
Ramsköld, L. 1986. Silurian encrinurid trilobites from Gotland and Dalarna, Sweden. Palaeontology, 29:527575.Google Scholar
Raymond, P. E. 1921. A contribution to the description of the fauna of the Trenton Group. Canada, Geological Survey, Museum Bulletin 31, 64 p.Google Scholar
Raymond, P. E., and Barton, D. C. 1913. A revision of the American species of Ceraurus. Bulletin of the Museum of Comparative Zoology at Harvard College, 54:523543.Google Scholar
Reed, F. R. C. 1899. The Lower Palaeozoic bedded rocks of County Waterford. The Quarterly Journal of the Geological Society of London, 55:718772.CrossRefGoogle Scholar
Reed, F. R. C. 1914. The Lower Palaeozoic trilobites of Girvan, Supplement. Palaeontographical Society, 1913, 56 p.Google Scholar
Reed, F. R. C. 1928. Notes on the family Encrinuridae. The Geological Magazine, 65:5177.CrossRefGoogle Scholar
Reed, F. R. C. 1931. The Lower Palaeozoic trilobites of Girvan, Supplement No. 2. Being a revision of some species previously described. Palaeontographical Society, 1929, 30 p.Google Scholar
Reed, F. R. C. 1935. The Lower Palaeozoic trilobites of Girvan, Supplement No. 3. Palaeontographical Society, 1934, 64 p.Google Scholar
Ride, W. D. L., Sabrosky, C. W., Bernardi, G., Melville, R. V., Corliss, J. O., Forest, J., Key, K. H. L., and Wright, C. W. 1985. International Code of Zoological Nomenclature. Third Edition. International Trust for Zoological Nomenclature, 338 p.Google Scholar
Ross, R. J. Jr., and Shaw, F. C. 1972. Distribution of the Middle Ordovician Copenhagen Formation and its trilobites in Nevada. (United States) Geological Survey Professional Paper 749, 33 p.CrossRefGoogle Scholar
Ross, R. J. Jr., and Shaw, F. C., et al. [27 authors]. 1982. The Ordovician System in the United States. Correlation chart and explanatory notes. International Union of Geological Sciences Publication, 12, 73 p.Google Scholar
Salter, J. W. 1848. In J. Phillips and Salter, Palaeontological Appendix to Professor John Phillips' Memoir on the Malvern Hills …. Memoir Geological Survey of Great Britain, 2(1) (fide Whittington, 1950).Google Scholar
Shaw, F. C. 1968. Early Middle Ordovician Chazy trilobites of New York. New York State Museum and Science Service, Memoir 17, 163 p.Google Scholar
Shaw, F. C. 1974. Simpson Group (Middle Ordovician) trilobites of Oklahoma. The Paleontological Society Memoir, 6, 54 p.Google Scholar
Shaw, F. C., and Lespérance, P. J. 1994. North American biogeography and taxonomy of Cryptolithus (Trilobita, Ordovician). Journal of Paleontology, 68:808823.CrossRefGoogle Scholar
Sinclair, G. W. 1947. Two examples of injury in Ordovician trilobites. American Journal of Science, 245:250257.CrossRefGoogle Scholar
Sinclair, G. W. 1964. Some Middle Ordovician fossils from Central Ontario, p. 3742. In Geology of Central Ontario. The American Association of Petroleum Geologists, Society of Economic Paleontologists and Mineralogists, Guidebook.Google Scholar
Slocom, A. W. 1913. New trilobites from the Maquoketa Beds of Fayette County, Iowa. Field Museum of Natural History, Publication 171, Geological Series, 4:4183.Google Scholar
Smith, A. B. 1994. Systematics and the Fossil Record: Documenting Evolutionary Patterns. Blackwell Scientific Publications, Oxford, 223 p.CrossRefGoogle Scholar
Stevens, P. F. 1991. Character states, morphological variation, and phylogenetic analysis: a review. Systematic Botany, 16:553583.CrossRefGoogle Scholar
Strusz, D. L. 1980. The Encrinuridae and related trilobite families, with a description of Silurian species from Southeastern Australia. Palaeontographica, Abteilung A, 168:168.Google Scholar
Sweet, W. C. 1984. Graphic correlation of upper Middle and Upper Ordovician rocks, North American Midcontinent Province. U.S.A., p. 2335. In Bruton, D. L. (ed.), Aspects of the Ordovician System. Palaeontological Contributions from the University of Oslo, 295, Universitetsforlaget.Google Scholar
Temple, J. T., and Tripp, R. P. 1979. An investigation of the Encrinurinae (Trilobita) by numerical taxonomic methods. Transactions of the Royal Society of Edinburgh, 70:223250.CrossRefGoogle Scholar
Temple, J. T., and Wu, Hong-Ji. 1990. Numerical taxonomy of Encrinurinae (Trilobita): additional species from China and elsewhere. Transactions of the Royal Society of Edinburgh: Earth Sciences, 81:209219.CrossRefGoogle Scholar
Tremblay, J. V., and Westrop, S. R. 1991. Middle Ordovician (Whiterockian) trilobites from the Sunblood Formation, District of Mackenzie, Canada. Journal of Paleontology, 65:801824.CrossRefGoogle Scholar
Tripp, R. P. 1962. Trilobites from the confinis flags (Ordovician) of the Girvan district, Ayrshire. Transactions of the Royal Society of Edinburgh, 65:140.CrossRefGoogle Scholar
Tripp, R. P. 1965. Trilobites from the Albany division (Ordovician) of the Girvan district, Ayrshire. Palaeontology, 8:577603.Google Scholar
Tripp, R. P. 1967. Trilobites from the Upper Stinchar Limestone (Ordovician) of the Girvan district, Ayrshire. Transactions of the Royal Society of Edinburgh, 67:4393.CrossRefGoogle Scholar
Tripp, R. P. 1974. New encrinurid trilobites from the Galena Formation (Ordovician) of Wisconsin and Iowa. Journal of Paleontology, 48:484488.Google Scholar
Tripp, R. P. 1979. Trilobites from the Ordovician Auchensoul and Stinchar Limestones of the Girvan District, Strathclyde. Palaeontology, 22:339361.Google Scholar
Tripp, R. P. 1980a. Trilobites from the Ordovician Balclatchie and lower Ardwell groups of the Girvan district, Scotland. Transactions of the Royal Society of Edinburgh: Earth Sciences, 71:123145.CrossRefGoogle Scholar
Tripp, R. P. 1980b. Trilobites from the Ordovician Ardwell Group of the Craighead Inlier, Girvan district, Scotland. Transactions of the Royal Society of Edinburgh: Earth Sciences, 71:147157.CrossRefGoogle Scholar
Troedsson, G. F. 1928. I. On the Middle and Upper Ordovician faunas of Northern Greenland. Part II. Jubilæumsekspeditionen Nord om Gr⊘nland 1920–23, number 7, 197 p.Google Scholar
Walcott, C. D. 1875. New species of trilobite from the Trenton Limestone at Trenton Falls, N. Y. Cincinnati Quarterly Journal of Science, 2:347349.Google Scholar
Walcott, C. D. 1877 [1879]. Description of new species of fossils from the Chazy and Trenton limestones, p. 6871. In Thirty-first Annual Report on the New York State Museum of Natural History by the Regents of the University of the State of New York, Albany [first published 1877: see p. 6 of report].Google Scholar
Westrop, S. R., and Ludvigsen, R. 1983. Systematics and paleoecology of Upper Ordovician trilobites from the Selkirk Member of the Red River Formation, southern Manitoba. Manitoba, Department of Energy and Mines, Mineral Resources Division, Geological Report GR 82–2, 51 p.Google Scholar
Whittington, H. B. 1941. Silicified Trenton trilobites. Journal of Paleontology, 15:492522.Google Scholar
Whittington, H. B. 1950. Sixteen Ordovician genotype trilobites. Journal of Paleontology, 24:531565.Google Scholar
Whittington, H. B. 1952. A unique remopleuridid trilobite. Museum of Comparative Zoology, Breviora Number 4, p. 19.Google Scholar
Whittington, H. B. 1954. Ordovician trilobites from Silliman's fossil mount, p. 119149. In Miller, A. K., Youngquist, W., and Collinson, C. (eds.), Ordovician Cephalopod Fauna of Baffin Island. Geological Society of America Memoir, 62.Google Scholar
Whittington, H. B. 1965. A monograph of the Ordovician trilobites of the Bala area, Merioneth, Part 2, p. 3362. Palaeontographical Society Monographs.CrossRefGoogle Scholar
Whittington, H. B. 1992. Fossils Illustrated. Volume 2. Trilobites. Boydell Press, 145 p., 120 plates.Google Scholar
Wiley, E. O., Siegel-Causey, D., Brooks, D. R., and Funk, V. A. 1991. The compleat cladist—a primer of phylogenetic procedures. The University of Kansas, Museum of Natural History, Special Publication No. 19, 158 p.CrossRefGoogle Scholar