Lingelsheimia
) are distinct from the rest of the subfamily in having a thick inner integument (over six cells thick), an exotegmen composed of cuboidal cells (type II), and vascular bundles in the outer integument and, as molecular evidence also suggests, should be transferred to a separate family Putranjivaceae. Hymenocardieae (Didymocistus and Hymenocardia), whose positions have been controversial, are monophyletic in sharing endotestal seeds with a collapsed exotegmen which is unknown elsewhere in Euphorbiaceae. The genera seem to require separation from the Euphorbiaceae. In addition, a morphological heterogeneity of the two large genera Cleistanthus and Phyllanthus, as well as of tribe Antidesmeae subtribe Scepinae were also discussed.
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Received 20 October 2000/ Accepted in revised form 14 January 2001
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Tokuoka, T., Tobe, H. Ovules and Seeds in Subfamily Phyllanthoideae (Euphorbiaceae): Structure and Systematic Implications. J Plant Res 114, 75–92 (2001). https://doi.org/10.1007/PL00013970
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DOI: https://doi.org/10.1007/PL00013970